Tiphiidae

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Tiphioidea

Familia: Tiphiidae
Subfamiliae (3): Brachycistidinae - Tiphiinae
Overview of genera

Acablasa Argaman, 1992
Acanthetropis Wasbauer, 1958
Aelurus Klug, 1840
Aeolothynnus Ashmead, 1903
Afromeria Boni Bartalucci, 2007
Aglyptacros Mickel & Krombein, 1942
Agriomyia Guérin-Méneville, 1838
Allenella Genise, 1992
Allomeria Boni Bartalucci, 2007
Ammodromus Guérin-Méneville, 1838
Anthobosca Guérin-Méneville, 1838
Anthosila Genise, 1985
Argenthynnus Genise, 1991
Arthrothynnus Brown, 1996
Aspidothynnus Turner, 1910
Astigmometopa Mickel & Krombein, 1942
Atopothynnus Kimsey, 1991
Aulacros Mickel & Krombein, 1942
Austromyzinum Brown, 1985
Beithynnus Kimsey, 2002
Bifidothynnus Brown, 1992
Brachymaya Kimsey & Wasbauer, 1999
Brethynnus Genise, 1991
Burdufa Argaman, 1992
Cabaraxa Nagy, 1974
Caetrathynnus Brown, 2000
Calchaquila Genise, 1985
Campylothynnus Turner, 1910
Catocheilus Guérin-Méneville, 1842
Chilothynnus Brown, 1996
Cleftomyzinum Brown, 1985
Colocistis Krombein, 1942
Cophothynnus Turner, 1908
Cosila Guérin-Méneville, 1838
Deuterothynnus Brown, 2005
Diamma Westwood, 1835
Dimorphothynnus Turner, 1910
Dolichetropis Wasbauer, 1968
Dorathynnus
Dryophia Antropov, 2014
Dythynnus Kimsey, 2001
Eirone Westwood, 1844
Elaphroptera Guérin-Méneville, 1838
Enteles Westwood, 1844
Epactiothynnus Turner, 1910
Epomidiopteron Romand, 1835
Eurycros Krombein, 1942
Foforoxia Argaman, 1992
Gebiupala Argaman, 1992
Glottynnus Genise, 1991
Glyptacros Mickel & Krombein, 1942
Guerinius Ashmead, 1903
Hadrocistis Wasbauer, 1968
Hathynnus Kimsey, 2003
Heligmothynnus Brown, 2005
Hemithynnus Ashmead, 1903
Hylomesa Krombein, 1968
Iswaroides Ashmead, 1899
Karlissa Krombein, 1979
Kaysara Carnimeo & Noll, 2018
Krombeinia Pate, 1947
Lalapa Pate, 1947
Lamprowara Boni Bartalucci, 2004
Leptothynnus Turner, 1910
Lestricothynnus Turner, 1910
Locodamia Argaman, 1992
Lophocheilus Guérin-Méneville, 1842
Ludita Nagy, 1967
Mallochia Allen, 1972
Megalothynnus Turner, 1910
Melaniswara Gorbatovskii, 1977
Meria Illiger, 1807
Meria Kimsey, 2011
Meriodes Boni Bartalucci, 2007
Merithynnus Kimsey, 1991
Mesa Kimsey, 2011
Mesa Saussure, 1890
Mesothynnus Kimsey, 1991
Methocha Kimsey, 2011
Methocha Latreille, 1804
Myrmosina Krombein, 1940
Myzinella Guiglia, 1959
Myzinum Latreille, 1803
Neotiphia Malloch, 1918
Neozeleboria Rohwer, 1910
Neozeloboria
Nitidothynnus Brown, 2000
Notomeria Boni Bartalucci, 2011
Ocasasla Argaman, 1992
Ornepetes Guérin-Méneville, 1838
Pampathynnus Carnimeo & Noll, 2018
Pandasaria Argaman, 1992
Paraquemaya Kimsey & Wasbauer, 1999
Paratiphia Saussure & Sichel, 1864
Pentazeleboria Brown, 1983
Philoponites Cockerell, 1915
Phymatothynnus Turner, 1908
Plesiomorpha Brèthes, 1913
Poecilotiphia Cameron, 1902
Pogonothynnus Turner, 1910
Procerothynnus Brown, 2000
Psammothynnus Ashmead, 1903
Pseudoscotaena Carnimeo & Noll, 2018
Pseudotiphia Ashmead, 1903
Pterombrus Smith, 1869
Punctotiphia Tsuneki, 1986
Rhagigaster Guérin-Méneville, 1838
Rhytidothynnus Brown, 2008
Rostrynnus Genise, 1991
Rugosothynnus
Sasmarila Argaman, 1992
Scotaena Klug, 1810
Sedomaya Kimsey & Wasbauer, 1999
Serpapinta Argaman, 1992
Sierocolpa Nagy, 1967
Silifka Argaman & Ozbek, 1992
Stilbopogon Mickel & Krombein, 1942
Tachynoides Kimsey, 1996
Tachynomyia Guérin-Méneville, 1842
Tachyphron Brown, 1995
Takyomyia Kimsey, 1996
Tamerlanella Boni Bartalucci, 2004
Telephoromyia Guérin-Méneville, 1838
Thanatotiphia Engel, Ortega-Blanco & Bennett, 2009
Thynnoides Guérin-Méneville, 1838
Tiphia J.C.Fabricius, 1775
Tiphia Kimsey, 2011
Tmesothynnus Turner, 1910
Umbothynnus Brown, 2008
Upa Kimsey, 1991
Vacacunda Argaman, 1992
Xeroglypta Mickel & Krombein, 1942
Zaspilothynnus Ashmead, 1903
Zaspolothynnus
Zeena Kimsey, 1991
Zeleboria Saussure, 1867
Zythynnus Kimsey, 2001
References

Bartalucci, M.B., 2009: Afrotropical species of the ancient genus Meria Illiger 1807 (Hymenoptera, Tiphiidae). Linzer Biologische Beiträge, 41 (2): 1817- 1861. Full article: [1]
Kimsey, L.S. 2011: Tiphiidae wasps of Madagascar (Hymenoptera, Tiphiidae). Journal of Hymenoptera research, 22: 45–68. DOI: 10.3897/JHR.22.1142
Lucia, M. et al. 2013: The identity of the enigmatic genus Stenocolletes Schrottky (Hymenoptera: Tiphiidae). Zootaxa 3652(4): 492–496. DOI: 10.11646/zootaxa.3652.4.7 Reference page.
Narita, K. & Mita, T. 2018. Two new species of the genus Methocha from Laos (Hymenoptera, Tiphiidae). ZooKeys 775: 59–68. DOI: 10.3897/zookeys.775.24945 Reference page.
Pilgrim, E.M.; von Dohlen, C.D.; Pitts, J.P. 2008: Molecular phylogenetics of Vespoidea indicate paraphyly of the superfamily and novel relationships of its component families and subfamilies. Zoologica scripta 37(5): 539–560. DOI: 10.1111/j.1463-6409.2008.00340.x Reference page.

Vernacular names
English: Tiphiid wasps
Nederlands: Keverdoders

The Tiphiidae (also known as the tiphiid wasps) are a family of large, solitary wasps whose larvae are parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea. Until recently, this family contained several additional subfamilies, but multiple studies have independently confirmed that these comprise a separate lineage, and are now classified in the family Thynnidae.[1][2]

The females of some Brachycistidinae are wingless, and hunt ground-dwelling (fossorial) beetle larvae. The prey is paralysed with the female's sting, and an egg is lain on it so the wasp larva has a ready supply of food. As some of the ground-dwelling scarab species attacked by tiphiids are pests, some of these wasps are considered beneficial as biological control agents.

Taxonomy

Tiphiid genera are classified as follows: [3][4][5]
The male of a species of Brachycistidinae photographed in Nevada
A female Tiphia femorata photographed in Italy
Subfamily Brachycistidinae

Acanthetropis Wasbauer, 1958
Brachycistellus Baker, 1907
Brachycistina Malloch, 1926
Brachycistis Fox, 1893
Brachymaya Kimsey & Wasbauer 1999
Colocistis Krombein, 1942
Dolichetropis Wasbauer, 1968
Glyptacros Mickel & Krombein, 1942
Hadrocistis Wasbauer, 1968
Paraquemaya Kimsey & Wasbauer, 1999
Sedomaya Kimsey & Wasbauer, 1999
Stilbopogon Mickel & Krombein, 1942

Subfamily Tiphiinae

Cabaraxa Nagy, 1974
Cyanotiphia Cameron, 1907
Epomidiopteron Romand, 1835
Icronatha Nagy, 1967
Krombeinia Pate, 1947
Ludita Nagy, 1967
Mallochessa Allen, 1972
Megatiphia Kimsey, 1993
Neotiphia Malloch, 1918
Paratiphia Sichel, 1864
Pseudotiphia Ashmead 1903
Tiphia Fabricius, 1775

Examples

Tiphia femorata Linden 1827
Tiphia minuta Fabricius, 1775

References

Pilgrim, E.; von Dohlen, C.; Pitts, J. (2008). "Molecular phylogenetics of Vespoidea indicate paraphyly of the superfamily and novel relationships of its component families and subfamilies". Zoologica Scripta. 37 (5): 539–560. doi:10.1111/j.1463-6409.2008.00340.x. S2CID 85905070.
Johnson, B.R.; et al. (2013). "Phylogenomics Resolves Evolutionary Relationships among Ants, Bees, and Wasps". Current Biology. 23 (20): 2058–2062. doi:10.1016/j.cub.2013.08.050. PMID 24094856.
"Taxonomy for Tiphiinae". insectoid.info. Retrieved 13 May 2017.
"Tiphiinae". Fauna Europaea. Retrieved 13 May 2017.

Lynn S. Kimsey; Marius Wasbauer (2006). "Phylogeny and Checklist of the Nocturnal Tiphiids of the Western Hemisphere (Hymenoptera: Tiphiidae: Brachycistidinae)". Journal of Hymenoptera Research. 15 (1): 9–25.