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Bembidion quadrimaculatum (Carabus quadriguttatus Fabricius )

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga

Familia: Carabidae
Subfamilia: Trechinae
Tribus: Bembidiini
Genus: Bembidion
Species: Bembidion quadrimaculatum

Bembidion quadrimaculatum (Linnaeus, 1760)

Lorenz, W. 2018. CarabCat: Global database of ground beetles (version Oct 2017). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P.M., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2018. Species 2000 & ITIS Catalogue of Life, 20th December 2017. Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on November 25, 2018]

Bembidion quadrimaculatum is a species of ground beetle of the family Carabidae.[1][2][3][4] It is found in Europe and Northern Asia (excluding China), North America, and Southern Asia.[1]


These six subspecies belong to the species Bembidion quadrimaculatum:

Bembidion quadrimaculatum caporiaccoi Netolitzky, 1935 c g
Bembidion quadrimaculatum cardiaderum Solsky, 1874 c g
Bembidion quadrimaculatum dubitans (LeConte, 1852) i c g b
Bembidion quadrimaculatum mandli Netolitzky, 1932 c g
Bembidion quadrimaculatum oppositum Say, 1823 i c g b
Bembidion quadrimaculatum quadrimaculatum (Linnaeus, 1761) i c g

Data sources: i = ITIS,[1] c = Catalogue of Life,[2] g = GBIF,[3] b =[4]

"Bembidion quadrimaculatum Report". Integrated Taxonomic Information System. Retrieved 2018-04-27.
"Bembidion quadrimaculatum species details". Catalogue of Life. Retrieved 2018-04-27.
"Bembidion quadrimaculatum". GBIF. Retrieved 2018-04-27.

"Bembidion quadrimaculatum Species Information". Retrieved 2018-04-27.

Further reading

Lindroth, Carl H. (1963). "The Ground-Beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska, part 3". Opuscula Entomologica Supplementum. 98 (2): 201–408.
Lobl, I.; Smetana, A., eds. (2017). Catalogue of Palaearctic Coleoptera, Volume 1: Archostemata - Myxophaga - Adephaga. Apollo Books. ISBN 978-90-04-33029-0.

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