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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Pyrginae
Tribus: Erynnini
Subtribus (2): Clitina – Erynnina

AnastrusAnaxasCamptopleuraChiomara – Chiothion – Clito – Crenda – CycloglyphaEbrietas – Echelatus – EphyriadesErynnis – Festivia – Gesta – Gorgythion – Helias – Hoodus – Mylon – Neomorphuncus – PotamanaxasSostrataTheagenesTimochares


Erynnini Brues & Melander, 1932: 236.

Brues, C. & Melander, A., 1932. Classification of Insects. Bulletin of the Museum of Comparative Zoology at Harvard College 73: 672pp., 1121 figs. BHL. Reference page.
Burns, J.M., Janzen, D.H., Hallwachs, W. & Grishin, N.V. 2019. Monotypy Justified: Genitalia and DNA Move Ephyriades eugramma (Mabille) to New Genus: Neomorphuncus Burns (Lepidoptera: Hesperiidae: Pyrginae). Proceedings of the Entomological Society of Washington 121(4): 557–567. DOI: 10.4289/0013-8797.121.4.557 Reference page.
Cong, Q., Zhang, J., Shen, J.H. & Grishin, N.V. 2019. Fifty new genera of Hesperiidae (Lepidoptera). Insecta Mundi 0731: 1–56. PDF. Reference page.
Li, W., Cong. Q., Shen, J.H., Zhang, J., Hallwachs, W., Janzen, D.H. & Grishin, N.V. 2019. Genomes of skipper butterflies reveal extensive convergence of wing patterns. PNAS 116(13): 6232–6237. DOI: 10.1073/pnas.1821304116. Text. SI Appendix. Reference page.

The Erynnini are a tribe in the skipper butterfly subfamily Pyrginae. They are a moderately diverse but quite plesiomorphic and inconspicuous group, and except for the Holarctic species of the type genus Erynnis occur only in the Neotropics.[1]

Formerly, when only four tribes of Pyrginae were recognized, they were included in the Pyrgini, which at that time contained a massive number of genera. But the Pyrginae have since been reorganized to make them and their tribes monophyletic, leading most modern authors to treat the Erynnini as distinct tribe. However, the old circumscription of the Pyrgini was by and large just as correct from a phylogenetic perspective.[1]

The tribe of the Pyrgini sensu lato most closely related to the Pyrgini sensu stricto are the strikingly different Achlyodidini. As many consider it desirable to treat this lineage as a distinct tribe, the Erynnini naturally need to be considered distinct too.[1]

The genera are listed here in the presumed phylogenetic sequence:[1]

Ephyriades Hübner, [1819]
Grais – hermit skipper
Erynnis – duskywings


Brower (2009)


Brower, Andrew V.Z. (2009): Tree of Life Web Project – Erynnini. Version of 2009-JUN-14. Retrieved 2009-DEC-26.

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