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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Cladus: Phytophaga
Superfamilia: Chrysomeloidea

Familia: Chrysomelidae
Subfamilia: Cryptocephalinae
Tribus: Fulcidacini
Genera: Aulacochlamys – Carcinobaena – ChlamisusDiplacaspis – Exema – Hymetes – Kakita – Melittochlamys – Neochlamisus – Poropleura – Pseudochlamys
[list of genera after Chamorro-Lacayo & Konstantinov (2009)]

Fulcidacini Jacobson, 1924

Type genus: Fulcidax Clavareau, 1913 [= Poropleura Lacordaire, 1848].

Chlamydeae Lacordaire, 1848: 636 [permanently invalid (Art. 39): based on preoccupied type genus]
Fulcidacina Jacobson, 1924: 239
Chlamisinae Gressitt, 1946: 84 [replacement name for Chlamydinae Lacordaire, 1848]
Chlamisini Gressitt, 1946

Primary references

Gressitt, J.L. 1946. Chinese chrysomelid beetles of the subfamily Chlamisinae. Annals of the Entomological Society of America 39(1): 84–100. DOI: 10.1093/aesa/39.1.84 Paywall.
Jacobson, G.G. 1924. Annotationes synonymicae et systematicae de Coleopteris. Russkoe Entomologicheskoe Obozrenie 18 [1922–1924] (4): 237–244. Full volume (PDF).
Lacordaire, J.T. 1848. Monographie des coléoptères subpentamères de la famille des phytophages. Tome second. Mémoires de la Société Royale des Sciences de Liège 5: vi + 890 pp. BHL [May 1848 (vol. title page)] Reference page.

Additional references

Doweld, A.B. 2014. Aulacomela, a new generic replacement name for Aulacochlamys Monrós 1952 (Insecta: Coleoptera: Chrysomelidae) non Margalef 1946 (Protista: Prasinomonada).Journal of Entomology and Zoology Studies 2(6): 299–300. PDF Free Access Reference page.
Agrain, F.A., Chamorro, M.L., Cabrera, N., Sassi, D. & Roig-Juñent, S. 2017. A comprehensive guide to the Argentinian case-bearer beetle fauna (Coleoptera, Chrysomelidae, Camptosomata). ZooKeys 677: 11–88. DOI: 10.3897/zookeys.677.10778 Open access. Reference page.
Bezděk, J. 2010. Cryptocephalinae: Fulcidacini. Pp. 617–619. In: Löbl, I. & Smetana, A. (eds.) Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Apollo Books, Stenstrup, Denmark. 924 pp. ISBN 978-87-88757-84-2. Google Books.
Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Campaner, C. 2014. Lista dos tipos da tribo Fulcidacini (Coleoptera, Chrysomelidae, Cryptocephalinae) depositados no Museu de Zoologia da Universidade de São Paulo. Papéis Avulsos de Zoologia 54(17): 209–215. DOI: 10.1590/0031-1049.2014.54.17 Open access.
Chamorro-Lacayo, M.L. & Konstantinov, A.S. 2009. Synopsis of warty leaf beetle genera of the world (Coleoptera, Chrysomelidae, Cryptocephalinae, Chlamisini). ZooKeys 8: 63–88. DOI: 10.3897/zookeys.8.90 Open access. Reference page.
Hernández-Sosa, L., Romero-Nápoles, J., Valdez-Carrasco, J.M., Carrillo-Sánchez, J.L., Ramírez-Alarcón, S., Ordóñez-Reséndiz, M.M., & Niño-Maldonado, S. 2019. Revisión genérica de Fulcidacini Jakobson, 1924 (Chrysomelidae, Cryptocephalinae) para México. Acta Zoológica Mexicana (n.s.) 35: 1–16. DOI: 10.21829/azm.2019.3502198 Open access. Reference page.
Karren, J.B. 1972. A revision of the subfamily Chlamisinae of America north of Mexico (Coleoptera: Chrysomelidae). The University of Kansas Science Bulletin 49: 875–988. BHL.
Riley, E.G., Clark, S.M. & Seeno, T.N. 2003. Catalog of the leaf beetles of America North of Mexico (Coleoptera: Megalopididae, Orsodacnidae and Chrysomelidae, excluding Bruchinae). Coleopterists Society, Special Publication No. 1. Sacramento, CA. 290 pp. ISBN 978-0-9726087-1-8. PDF. Reference page.

The Fulcidacini are a tribe within the leaf beetle subfamily Cryptocephalinae,[1] though historically they were often treated as a distinct subfamily, Chlamisinae. 11 genera with altogether about 400 species are currently placed here; some four-fifths of the species are found in the Neotropics, but the rest is distributed over all other continents except Antarctica.[2]

Genera and some selected species include:

Chlamisus Rafinesque, 1815
Chlamisus amyemae Reid, 1991
Chlamisus arizonensis Linell, 1898
Chlamisus aterrimus Lea, 1904
Chlamisus flavidus Karren, 1972
Chlamisus foveolatus Knoch, 1801
Chlamisus huachucae Schaeffer, 1906
Chlamisus maculipes Chevrolat, 1835
Chlamisus mimosae Karren, 1989
Chlamisus minax Lacordaire
Chlamisus nigromaculatus Karren, 1972
Chlamisus quadrilobatus Schaeffer, 1926
Chlamisus texanus Schaeffer, 1906
Diplacaspis Jacobson, 1924
Diplacaspis prosternalis Schaeffer, 1906
Poropleura Lacordaire, 1848 (=Fulcidax Voet, 1806, an unavailable name[3])
Poropleura bacca (Kirby, 1818)
Poropleura coelestina (Lacordaire, 1848)
Poropleura monstrosa (Fabricius, 1798)
Melitochlamys Monrós, 1948
Neochlamisus – warty leaf beetles (c.17 species)
Pseudochlamys Lacordaire, 1848
Pseudochlamys megalostomoides Lacordaire, 1849
Pseudochlamys semirufescens Karren, 1972


Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Ślipiński, A.; Smith, A. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
Chaboo et al. (2008)

Krell, F-T. (2012) On nomenclature and synonymy of Trichius rosaceus, T. gallicus, and T. zonatus (Coleoptera: Scarabaeidae: Cetoniinae: Trichiini). Zootaxa 3278: 61–68.


Chaboo, Caroline S.; Brown, Christopher G. & Funk, Daniel J. (2008): Faecal case architecture in the gibbosus species group of Neochlamisus Karren, 1972 (Coleoptera: Chrysomelidae: Cryptocephalinae: Chlamisini). Zool. J. Linn. Soc. 152(2): 315–351. doi:10.1111/j.1096-3642.2007.00343.x (HTML abstract)

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