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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Paraneoptera
Superordo: Condylognatha
Ordo: Hemiptera
Subordo: Heteroptera
Infraordo: Pentatomomorpha
Superfamilia: Pentatomoidea

Familia: Pentatomidae
Subfamilia: Podopinae
Tribus: Graphosomatini
Genus: Graphosoma
Subgenera: G. (Graphosoma) – G. (Graphosomella)

Species: G. alkani – G. consimile – G. inexpectatum – G. italicum – G. interruptum – G. lineatum – G. melanoxanthum – G. rubrolineatum – G. semipunctatum – G. stali

Graphosoma de Laporte, 1832

Type species: Cimex nigrolineatus Fabricius, 1781 [=Cimex lineatus Linnaeus, 1758, by subsequent designation Reuter, 1888.]

Primary references

Laporte, F.L. 1833. Essai d'une classification systématique de l'ordre des Hémiptères (Hémiptères Héteroptères, Latr.). Magasin de Zoologie 2: 1–88. BHL Reference page. [p. 70]
Reuter, O.M. 1888. Revision synonymica heteropterorum palaearcticorum quae descripserunt auctores vetustiores (Linnaeus 1758—Latreille 1806). Helsingfors [Helsinki]: 1–485. [also published in Acta Societatis Scientiarum Fennica 15(1888): 443–812. BHL Reference page. [p. 461–462]

Additional references

Carapezza, A. & Jindra, Z. 2008. Graphosoma (Graphosomella subgen. nov.) inexpectatum sp. nov. from Syria (Hemiptera Heteroptera Pentatomidae). Naturalista Siciliano (4), 32(3–4): 471–478. PDF Reference page.
Lupoli, R. & Dusoulier, F. 2015. Les Punaises Pentatomoidea de France pp.429. Éditions Ancyrosoma. [In French]. ISBN 978-2-9536661-1-3. ResearchGate Paywall Reference page.
Lupoli, R. 2017. Graphosoma lineatum (L., 1758) and G. italicum (O.F. Müller, 1766), two valid and distinct species, probably derived from the Zanclean mediterranean transgression (Hemiptera Pentatomidae). L'Entomologiste 73(1): 19–33. [In French]. ResearchGate Open access. Reference page.
Péricart, J. 2010. Hémiptères Pentatomoidea Euro-Méditerranéens. Volume 3 : Podopinae et Asopinae. Faune de France 93: 1–290. Fédération française des sociétés de sciences naturelles, 290 p. ISBN 978-2-903052-32-4 [In French]. Preview. Reference page. (p. 43–48)
Rider, D.A. 2006. Family Pentatomidae. p. 233–403, In: Aukema, A. & Rieger, C. (eds). Catalogue of the Heteroptera of the Palaearctic Region, vol 5. The Netherlands Entomological Society, Amsterdam. ResearchGate Open access PDF. Reference page.


Graphosoma Laporte de Castelnau, 1833 – Taxon details on Biological Library (BioLib).
Graphosoma Laporte 1833 – Taxon details on Encyclopedia of Life (EOL).
Graphosoma Laporte, 1833 – Taxon details on Fauna Europaea.
Graphosoma – Taxon details on Global Biodiversity Information Facility (GBIF).
Graphosoma - Taxon details on Index to Organism Names (ION).

Graphosoma – Taxon details on National Center for Biotechnology Information (NCBI).
Graphosoma – Taxon details on Nomenclator Zoologicus (NZ).
Graphosoma - Taxon details on Pentatomoidea Classification accessed on 06/11/2019

Vernacular names
magyar: Pajzsos poloska

Graphosoma is a genus of the family Pentatomidae known as the striped shield bugs for their distinctive markings.
Subgenus Graphosoma

Graphosoma alkani Lodos, 1959
Graphosoma consimile Horváth, 1903
Graphosoma interruptum White, 1839
Graphosoma italicum (Müller, 1766)
Graphosoma lineatum (Linnaeus, 1758)
Graphosoma melanoxanthum Horváth, 1903
Graphosoma rubrolineatum (Westwood, 1837)
Graphosoma semipunctatum (Fabricius, 1775)
Graphosoma stali Horváth, 1881

Subgenus Graphosomella

Graphosoma inexpectatum Carapezza & Jindra, 2008


Carapezza, A.; Jindra, Z. 2008: Graphosoma (Graphosomella subgen. nov.) inexpectatum sp. nov. from Syria (Hemiptera Heteroptera Pentatomidae). Naturalista siciliano (4), 32(3-4): 471–478. Full article (PDF)
Dai, J.-x. 2005: Discussion on classification of genus Grophosoma [Graphosoma] and Dybowskyia based on sequences of cytochrome b gene (Hemiptera: Pentatomidae). Sichuan journal of zoology, 24(4): 490–495.
Fent, M.; Dursun, A.; Tezcan, S. 2013: First record of Graphosoma inexpectatum (Hemiptera, Pentatomidae, Podopinae) from Turkey with description of the female. ZooKeys, 319: 51–57. doi:10.3897/zookeys.319.4298

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