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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Coccinelloidea

Familia: Coccinellidae
Subfamilia: Coccinellinae
Tribus: Coccinellini
Genus: Harmonia
Species (20): H. antipodum – H. axyridis – H. bicolor – H. conformis – H. coryphaea – H. dimidiata – H. eucharis – H. expalliata – H. lapeyrousei – H. manillana – H. octomaculata – H. pardalina – H. quadripunctata – H. sedecimnotata – H. shoichii – H. testudinaria – H. yedoensis – ...

Harmonia Mulsant, 1846
Type species: Coccinella marginepunctata Schaller, 1783 (= Coccinella quadripunctata Pontoppidan, 1763) by subsequent designation of Timberlake, 1943


Ballia Mulsant, 1853: 162
Callineda Crotch, 1871: 6 (syn. Timberlake, 1943: 17)
Leis Mulsant, 1850: 241 (syn. Miyatake, 1965: 60)
Ptychanatis Crotch, 1874: 122 (syn. Timberlake, 1943: 17)
Rhopaloneda Timberlake, 1943: 17n.
Stictoleis Crotch, 1874: 118

Primary references

Mulsant, E. 1846. Histoire Naturelle des Coléoptères de France. Sulcicolles-Sécuripalpes. Paris: Maison xxiv + 26 pp. + 280 pp. + 1 pl. Full text Google Books Reference page.
Poorani, J. & Booth, R.G. 2016. Harmonia manillana (Mulsant), a new addition to Indian Coccinellidae, with changes in synonymy. Biodiversity Data Journal 4: e8030. DOI: 10.3897/BDJ.4.e8030. Reference page.

Additional references

Crotch, G.R. 1871. List of Coccinellidae. Cambridge. 8 pp. Reference page.
Crotch, G.R. 1874. A revision of the coleopterous family Coccinellidae. E. W. Janson, London. xv. + 311 pp.Internet Archive BHL. Reference page.
Miyatake, M. 1965. Some Coccinellidae (excluding Scymnini) of Formosa (Coleoptera). Special Bulletin of the Lepidopterists’ Society of Japan, 1: 50–74. Full text Reference page.
Mulsant, E. 1853. Supplement a la monographie des Coléoptères trimères sécuripalpes. Annales de la Société Linnéenne de Lyon (2) 1: 129–333. BHL Reference page.
Timberlake P.H. 1943. The Coccinellidae or ladybeetles of the Koebele collection–Part I. Hawaiian Planters' Record 47: 1–67. Internet Archive Reference page.

Vernacular names
日本語: テントウムシ属

Harmonia is a genus of lady beetles belonging to the subfamily Coccinellinae.

The distinction of the individual species within the genus is often very difficult, and in some cases requires examination of the reproductive system. The coloring and drawing in the genus Harmonia are very variable, not only between species, but also within each species. The basic color of the elytra is usually red to orange with black spots. The patches can be linked together to form black bars. The species can reach a length of 3 to 7 mm. The rear edge of the pronotum is slightly curved. The antennae are a little shorter than the head and are composed of eleven segments. The three last segments are thickened at the tip and form a club.
List of species

Harmonia axyridis (Pallas, 1771) – Asian lady beetle, originally from Nepal, China, Taiwan, Japan, introduced to Western Europe and America
Harmonia antipodum (Mulsant, 1848) – antipodean ladybird, endemic to New Zealand
Harmonia basinotata Bielawski, 1964 – New Guinea
Harmonia bicolor (Blackburn, 1892) – Australia
Harmonia conformis (Boisduval, 1835) – common spotted lady beetle, Australia, introduced in New Zealand
Harmonia dimidiata (Fabricius, 1781) – India, Pakistan, Nepal, Bhutan, China, Taiwan, Japan; introduced in North America
Harmonia dunlopi (Crotch, 1874) – India, Malaysia, Philippines
Harmonia eucharis (Mulsant, 1850) – India, Pakistan, Myanmar, Nepal, Himalayan Region, South China
Harmonia expallida (Weise, 1907) – India
Harmonia octomaculata (Fabricius, 1781) – India, Pakistan, Nepal, Bangladesh, Sri Lanka, Micronesia, Australia
Harmonia quadripunctata (Pontoppidan, 1763) – cream-streaked lady beetle, Europe
Harmonia sedecimnotata (Fabricius, 1801) – India, Nepal, Southeast Asia
Harmonia testudinaria (Mulsant, 1850) – Indonesia, New Guinea, North Australia
Harmonia uninotata Bielawski, 1964 – New Guinea
Harmonia yedoensis (Takizawa, 1917) – Japan

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