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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Tenebrionoidea

Familia: Tenebrionidae
Subfamilia: Tenebrioninae
Tribus: Heleini
Subtribus: Cyphaleina

Genera: CelibeEmcephalus –Helea – Mimopeus


Watt, J.C. 1968: Specific synonymy in Mimopeus Pascoe (Cilibe auctorum), and the nomenclatural status of some related genera (Coleoptera: Tenebrionidae). New Zealand entomologist, 4(1): 35–39. BUGZ

Heleini is a tribe of darkling beetles in the family Tenebrionidae. There are more than 40 genera in Heleini, found in Australasia.[1][2]

These genera belong to the tribe Heleini:

Agastenes R. Lucas, 1920
Aglypta Gebien, 1908
Amarygmimus Bates, 1873
Amphianax Bates, 1873
Asphalus Pascoe, 1868
Atoreuma Gebien, 1941
Bassianus Matthews & Doyen, 1989
Bolbophanes Carter, 1913
Boreosaragus Matthews, 1993
Brises Pascoe, 1869
Byallius Pascoe, 1869
Camponotiphilus Lea, 1914
Celibe Boisduval, 1835
Cerodolus Sharp, 1886
Cillibus Matthews, 1993
Cyphaleus Westwood, 1841
Dysarchus Pascoe, 1866
Edylius Champion, 1894
Emcephalus W. Kirby, 1828
Helea Latreille, 1804
Hemicyclus Westwood, 1841
Meneristes Pascoe, 1869
Mimopeus Pascoe, 1866
Mithippia Pascoe, 1869
Mitrothorax Carter, 1914
Nyctozoilus Guérin-Méneville, 1831
Olisthaena Erichson, 1842
Onotrichus Carter, 1911
Ospidus Pascoe, 1866
Pachycoelia Boisduval, 1835
Paraphanes W.J. MacLeay, 1887
Phanechloros Matthews & Bouchard, 2008
Platyphanes Westwood, 1849
Prophanes Westwood, 1849
Pseudhelops Guérin-Méneville, 1841
Pterohelaeus Brême, 1842
Saragus Erichson, 1842
Sloanea Carter, 1916
Styrus Bates, 1873
Sympetes Pascoe, 1866
Trichosaragus Blackburn, 1890


Bouchard, Patrice; Bousquet, Yves; Aalbu, Rolf L.; Alonso-Zarazaga, Miguel A.; et al. (2021). "Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera)". ZooKeys (1050): 1–633. doi:10.3897/zookeys.1050.64217. hdl:10261/250214. PMC 8328949. PMID 34385881.

Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.

Further reading

LeConte, J. L. (1861). Classification of the Coleoptera of North America. Smithsonian Miscellaneous Collections. Vol. 3. Smithsonian Institution. doi:10.5962/bhl.title.38459. ISBN 978-0665100550.
Somerby, Ronald E.; Thomas, Donald B.; Triplehorn, Charles A. (2002). "Family 106: Tenebrionidae". In Arnett, Ross H. Jr.; Thomas, Michael C.; Skelley, Paul E.; Frank, J. Howard (eds.). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. pp. 463–509. ISBN 978-0-8493-0954-0.

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