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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Tenebrionoidea

Familia: Tenebrionidae
Subfamilia: Tenebrioninae
Tribus: Tenebrionini
Genus: Tenebrio
Subgenera: T. (Afrotenebrio) – T. (Tenebrio)

Species: T. alternicostis – T. angustus – T. cedrorum – T. fuliginosus – T. medius – – T. opacus – T. punctatissimus – T. punctipennis – T. syriacus – T. tristisT. molitor T. obscurus

Nomina dubia:

T. angulatus – T. barbarus – T. erraticus – T. pedicularius

Tenebrio Linnaeus, 1758

Type species: Tenebrio molitor Linnaeus, 1758

Primary references

Linnaeus, C. 1758. Systema Naturae per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Tomus I. Editio decima, reformata. Holmiæ: impensis direct. Laurentii Salvii. i–ii, 1–824 pp DOI: 10.5962/bhl.title.542: 417. Reference page.

Additional references

Ferrer, J.; Holston, K. 2009: Identities of Tenebrio Linnaeus types at Uppsala, and the resulting changes in old darkling beetle names (Insecta: Coleoptera: Tenebrionidae). Zootaxa, 2308: 29–42. Abstract & excerpt [erratum in Zootaxa, 2359: 68 (2010) PDF


Tenebrio Taxon details on Fauna Europaea
Atlas of darkling beetles (Tenebrionidae) of Russia
ZooBank: 1F3AF844-1E30-4943-A38F-1B740892812C

Vernacular names
suomi: Jauhopukit
հայերեն: Ալիւրաճճի
русский: Хрущак мучной

Tenebrio is a genus of darkling beetles.[2][3] Adults are 12–18 mm (0.47–0.71 in) long and can live for 1–2 years. The larvae are minor pests, but they are also widely reared and sold as pet food.[3]

The genus contains the following extant species:[2][3]

Tenebrio molitor Linnaeus, 1758 – yellow mealworm
Tenebrio obscurus Fabricius, 1792 – dark mealworm

Four species are known from fossils found in Germany and Canada:[1][4]

Tenebrio calculensis Scudder, 1895 - (Pleistocene, Leda Clay, Canada)
Tenebrio effossus Germar, 1837 - (Oligocene, Rott Formation, Germany)
Tenebrio primigenius Scudder, 1879 - (Ypresian, Allenby Formation, Canada)
Tenebrio senex Von Heyden, 1859 - (Oligocene, Rott Formation, Germany)


"†Tenebrio Linnaeus 1758". Paleobiology Database. Fossilworks. Retrieved 17 December 2021.
"Tenebrio". Integrated Taxonomic Information System. Retrieved 4 January 2017.
Rees, David (21 July 2004). Insects of Stored Products. CSIRO Publishing. pp. 112–113. ISBN 978-0-643-10263-7.
Scudder, S. H (1895). "Canadian fossil insects, myriapods and arachnids, Vol II. The Coleoptera hitherto found fossil in Canada". Geological Survey of Canada Contributions to Canadian Palaeontology. 2: 5–26.

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