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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Paraneoptera
Superordo: Condylognatha
Ordo: Hemiptera
Subordo: Heteroptera
Infraordo: Pentatomomorpha
Superfamilia: Pentatomoidea

Familia: Urostylididae
Subfamiliae (1): Urostylidinae
Overview of genera

Chelurotropella – Cobbenicoris – Eurhynchiocoris – Tessaromerus – Urochela – Urochelellus – Urolabida – Urostylis


The former spelling, "Urostylidae", is grammatically incorrect and improperly formed under ICZN Article 29.[1]

H. Berger, E. Heiss, I. M. Kerzhner (2001) Removal of Homonymy between Urostylidae DALLAS, 1851 (Insecta, Heteroptera) and Urostylidae BÜTSCHLI, 1889 (Ciliophora, Hypotrichia) Ann. Naturhist. Mus. Wien 103(B): 301-302.

Ahmad, I.; Moizuddin, M. 1992: A review and cladistics of Urostylidae Dallas (Hemiptera: Pentatomoidea) with keys to taxa of Indian subregion and description of four genera and five species including two new ones from Pakistan, Azad Kashmir and Bangladesh. Philippine journal of science, 121(3): 263–297. [not seen]
Kim, J., Roca-Cusachs, M. & Jung, S. 2018. Taxonomic review of the genus Urostylis (Hemiptera: Heteroptera: Urostylididae) from the Korean Peninsula, with description of a new species. Zootaxa 4433(3): 445–456. DOI: 10.11646/zootaxa.4433.3.3 Paywall Reference page.
Zhou, Y-Y. & Rédei, D. 2018. A new synonymy in East Asian Urostylididae (Hemiptera: Heteroptera). Zootaxa 4504(1): 145–150. DOI: 10.11646/zootaxa.4504.1.10 Paywall Reference page.

Urostylididae is a family of true bugs and is considered a basal or "primitive" family within the stink-bug lineage. They are found only in Asia. Older works used the spelling Urostylidae but this clashes with the name used for a protozoan family and a spelling correction (emendation) has been suggested that also avoids the confusion created by homonyms. The family name Urolabididae has also been used for some members in the past.[1]
Urolabida grayi

This family has about 100 species, with distribution limited mainly to eastern Asia. Urostylidids are 4 to 15 mm and somewhat longer than broad with elongated legs, three tarsal segments, and a small head. The antenna has 5 segments of which the first is longer than the head. The base of the antenna has broad tubercles and has a ringed appearance. The simple eyes or ocelli, when present (Urolabida lacks ocelli[2]), are very close to each other. They show a wide separation between the hind and mid coxae on the underside. Other important family characters are the presence of a structure for stridulation, the stridulitrum on the first anal vein of the hind wing, and bristles on the claws. The female genital structure has nine gonocoxites that form a M or W-shaped sclerite.[3] They suck plant sap.[4][5] The male reproductive anatomy suggests close affinities to the Tessaratomidae subfamily Natalicolinae. The spermatheca has two pump flanges in two genera which is a feature found also in the Aradidae and Leptopodidae. Thus the phylogeny of the group is unclear but lies within the Pentatomoidea.[6] In the past the family was divided into two subfamilies, but one group, the Saileriolinae, has since been suggested as a distinct family, Saileriolidae, by at least two studies. The position of Saileriolidae within the Pentatomoidea is still unclear.[3][7]

Berger, H.; Heiss, E.; Kerzhner, I.M. (2001). "Removal of homonymy between Urostylidae Dallas, 1851 (Insecta, Heteroptera) and Urostylidae Buetschli, 1889 (Ciliophora, Hypotrichia)" (PDF). Annalen des Naturhistorischen Museums in Wien. 103B: 301–302.
Distant, WL (1902). Fauna of British India. Rhynchota. Volume 1. London: Taylor and Francis. p. 303.
Grazia, Jocelia; Schuh, Randall T.; Wheeler, Ward C. (2008). "Phylogenetic relationships of family groups in Pentatomoidea based on morphology and DNA sequences (Insecta: Heteroptera)" (PDF). Cladistics. 24 (6): 932–976. doi:10.1111/j.1096-0031.2008.00224.x. PMID 34892882. S2CID 41951432.
Schuh, Randall T.; Slater, James Alexander (1995). True Bugs of the World (Hemiptera:Heteroptera): Classification and Natural History. Cornell University Press. pp. 245–246.
Ren, Shu-Zhi; Lin, Cheng-Shing (2003). "Revision of the Urostylidae of Taiwan, with descriptions of three new species and one new record (Hemiptera-Heteroptera: Urostylidae)" (PDF). Formosan Entomologist. 23: 129–143.
Kumar, R. (1971). "Morphology and relationships of the Pentatomoidea (Heteroptera) 5-Urostylidae". American Midland Naturalist. 85 (1): 63–73. doi:10.2307/2423912. JSTOR 2423912.
Yao, Yunzhi; Cai, Wanzhi; Rider, David A.; Ren, Dong (2012). "Primipentatomidae fam. nov. (Hemiptera: Heteroptera: Pentatomomorpha), an extinct insect family from the Cretaceous of north-eastern China". Journal of Systematic Palaeontology. 11: 1–20. doi:10.1080/14772019.2011.639814. S2CID 86753502.

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