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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Antliophora
Ordo: Diptera
Subordo: Brachycera
Infraordo: Stratiomyomorpha

Familia: †Zhangsolvidae

Arillo, A. et al. 2015: Long-proboscid brachyceran flies in Cretaceous amber (Diptera: Stratiomyomorpha: Zhangsolvidae). Systematic entomology, 40(1): 242–267. DOI: 10.1111/syen.12106 Reference page.

Zhangsolvidae is an extinct family of brachyceran flies known from the Cretaceous period. Members of the family possess a long proboscis, varying in length between 1.3 and 7 mm depending on the species, and were probably nectarivores. A specimen has been found with preserved Bennettitales pollen, suggesting that they acted as pollinators for extinct gymnosperms.[1] They are considered to be members of the Stratiomyomorpha.[2]

†Buccinatormyia Arillo et al. 2015[2]
†Buccinatormyia gangnami Khramov and Nam 2019[3] Jinju Formation , South Korea, Albian
†Buccinatormyia magnifica Arillo et al. 2015 Spanish amber, Albian
†Buccinatormyia soplaensis Arillo et al. 2015 Spanish amber, Albian
†Burmomyia Zhang and Wang 2019[4]
†Burmomyia rossi Zhang and Wang 2019 Burmese amber, Myanmar, Cenomanian
†Cratomyia Mazzarolo and Amorim 2000[5]
†Cratomyia cretacica Wilkommen 2007[6] Crato Formation , Brazil, Aptian
†Cratomyia macrorrhyncha Mazzarolo and Amorim 2000 Crato Formation, Brazil, Aptian
†Cratomyia mimetica Grimaldi 2016[7] Burmese amber, Myanmar, Cenomanian
†Cratomyia zhuoi Zhang and Wang 2019[4] Burmese amber, Myanmar, Cenomanian
†Linguatormyia Grimaldi 2015 in Arillo et al. 2015[2]
†Linguatormyia teletacta Grimaldi 2015 in Arillo et al. 2015 Burmese amber, Myanmar, Cenomanian
†Zhangsolva Nagatomi and Yang 1998[8]
†Zhangsolva cupressa Zhang et al. 1993[9] Laiyang Formation, China, Aptian
†Zhangsolva burmensis Zhang & Zhang, 2021[10] Burmese amber, Myanmar, Cenomanian


Peñalver, Enrique; Arillo, Antonio; Pérez-de la Fuente, Ricardo; Riccio, Mark L.; Delclòs, Xavier; Barrón, Eduardo; Grimaldi, David A. (July 2015). "Long-Proboscid Flies as Pollinators of Cretaceous Gymnosperms". Current Biology. 25 (14): 1917–1923. doi:10.1016/j.cub.2015.05.062. PMID 26166781. S2CID 13022302.
Arillo, Antonio; Peñalver, Enrique; Pérez-De La Fuente, Ricardo; Delclòs, Xavier; Criscione, Julia; Barden, Phillip M.; Riccio, Mark L.; Grimaldi, David A. (January 2015). "Long-proboscid brachyceran flies in Cretaceous amber (Diptera: Stratiomyomorpha: Zhangsolvidae): Cretaceous brachyceran family Zhangsolvidae". Systematic Entomology. 40 (1): 242–267. doi:10.1111/syen.12106. hdl:2445/59152. S2CID 82832797.
Khramov, Alexander V.; Nam, Gi-Soo; Vasilenko, Dmitry V. (2020-01-02). "First long-proboscid flies (Diptera: Zhangsolvidae) from the Lower Cretaceous of South Korea". Alcheringa: An Australasian Journal of Palaeontology. 44 (1): 160–168. doi:10.1080/03115518.2019.1664634. ISSN 0311-5518. S2CID 213651448.
Zhang, Qingqing; Chen, Keyu; Wang, Yating; Xue, Ruixuan; Jarzembowski, Edmund A.; Wang, Bo (June 2019). "Long-proboscid zhangsolvid flies in mid-Cretaceous Burmese amber (Diptera: Stratiomyomorpha)". Cretaceous Research. 98: 18–25. doi:10.1016/j.cretres.2019.01.019.
L. A. Mazzarolo and D. S. Amorim. 2000. Cratomyia macrorrhyncha, a Lower Cretaceous brachyceran fossil from the Santana Formation, Brazil, representing a new species, genus and family of the Stratiomyiomorpha (Diptera). Insect Systematics & Evolution 31:91-102
J. Wilkommen. 2007. Taxonomic names, in Diptera: true flies, gnats, and crane flies. The Crato Fossil Beds of Brazil: Window into an Ancient World 369-387
D. A. Grimaldi. 2016. Diverse orthorrhaphan flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History 408:1-131
A. Nagatomi and D. Yang. 1998. A review of extinct Mesozoic genera and families of Brachycera (Insecta, Diptera, Orthorrhapha). Entomologist's Monthly Magazine 134:95-192
J. F. Zhang, S. Zhang, and L. Y. Li. 1993. Mesozoic gadflies (Insecta: Diptera). Acta Palaeontologica Sinica 32:662-672
Zhang, Qingqing; Zhang, Junfeng (2021-12-07). "New record of Zhangsolva Nagatomi & Yang, 1998 (Diptera, Zhangsolvidae) from mid-Cretaceous Burmese amber and its implications". Geological Society, London, Special Publications. 521. doi:10.1144/SP521-2021-150. ISSN 0305-8719.

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