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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Crustacea
Superclassis: Multicrustacea
Classis: Malacostraca
Subclassis: Eumalacostraca
Superordo: Eucarida
Ordo: Decapoda
Subordo: Pleocyemata
Infraordo: Brachyura
Sectio: Eubrachyura
Subsectio: Thoracotremata
Superfamilia: Grapsoidea

Familia: Sesarmidae
Genus: Geosesarma
Species (53): G. aedituens – G. albomita – G. ambawang – G. amphinome – G. anambas – G. angustifrons – G. araneum – G. aurantium – G. batak – G. bau – G. bicolor – G. cataracta – G. celebense – G. cikaniki – G. clavicrure – G. confertum – G. danumense – G. foxi – G. garutense – G. gordonae – G. gracillimum – G. hednon – G. ianthina – G. insulare – G. johnsoni – G. katibas – G. krathing – G. larsi – G. lawrencei – G. lebak – G. leprosum – G. maculatum – G. malayanum – G. mirum – G. nannophyes – G. nemesis – G. noduliferum – G. notophorum – G. ocypodum – G. penangense – G. peraccae – G. pontianak – G. protos – G. pylaemenes – G. rathbunae – G. robustum – G. rouxi – G. sabanum – G. sarawakense – G. scandens – G. sekop – G. serenei – G. sodalis – G. solomonense – G. spectrum – G. starmuhlneri – G. sumatraense – G. sukabumi – G. sylvicola – G. ternatense – G. teschi – G. thelxinoe – G. tiomanicum – G. vicentense
Name

Geosesarma de Man, 1892

Type species:
References

de Man 1892 in Weber, Ergeb. Reise Niederl. O.-Ind., 2 (2), 341.
Ng, P.K.L. 2015. Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species. Zootaxa 4048(1): 37–56. DOI: 10.11646/zootaxa.4048.1.2. Preview (PDF) Reference page.
Ng, P.K.L. 2021. Geosesarma sodalis, a new species of vampire crab (Crustacea, Brachyura, Sesarmidae) from a limestone cave in central Sarawak, Malaysia. Zookeys 1031: 133–141. DOI: 10.3897/zookeys.1031.63134 Open access Reference page.
Ng, P.K.L. & Grinang, J. 2018. A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa 4508(4): 569–578. DOI: 10.11646/zootaxa.4508.4.5 Paywall Reference page.
Ng, P.K.L. & Lemaitre, R. 2017. On the taxonomy of two poorly known species of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) from the Philippines. Zootaxa 4250(4): 475–483. DOI: 10.11646/zootaxa.4250.4.5. Reference page.
Ng, Paul Y.C. & Ng, P.K.L. 2019. Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo. Zootaxa 4614(3): 529–540. DOI: 10.11646/zootaxa.4614.3.6 Paywall Reference page.
Shy, J-Y. & Ng, P.K.L. 2019. Geosesarma mirum, a new species of semi-terrestrial sesarmid crab (Crustacea, Decapoda, Brachyura) from central Taiwan. Zookeys 858: 1–10. DOI: 10.3897/zookeys.858.35198 Reference page.

Geosesarma is genus of small freshwater or terrestrial crabs, typically less than 10 mm (0.4 in) across the carapace.[2] They live and reproduce on land with the larval stages inside the egg. They are found from India,[3] through Southeast Asia, to the Solomon Islands and Hawaii.[2]

In the pet trade, they are sometimes called vampire crabs. This has nothing to do with their feeding habits, but rather with the bright, contrastingly yellow eyes of some Geosesarma species.[4]
Species

Geosesarma contains these species:[1]

Geosesarma aedituens Naruse & Jaafar, 2009
Geosesarma albomita Yeo & Ng, 1999
Geosesarma amphinome (De Man, 1899)
Geosesarma angustifrons (A. Milne-Edwards, 1869)
Geosesarma araneum (Nobili, 1899)
Geosesarma aurantium Ng, 1995
Geosesarma bau Ng & Jongkar, 2004
Geosesarma bicolor Ng & Davie, 1995
Geosesarma bintan T. M. Leong, 2014
Geosesarma cataracta Ng, 1986
Geosesarma celebense (Schenkel, 1902)
Geosesarma clavicrure (Schenkel, 1902)
Geosesarma confertum (Ortmann, 1894)
Geosesarma danumense Ng, 2003
Geosesarma dennerle Ng, Schubart & Lukhaup, 2015
Geosesarma foxi (Kemp, 1918)
Geosesarma gordonae (Serène, 1968)
Geosesarma gracillimum (De Man, 1902)
Geosesarma hagen Ng, Schubart & Lukhaup, 2015
Geosesarma hednon Ng, Liu & Schubart, 2003
Geosesarma ianthina Pretzmann, 1985
Geosesarma insulare Ng, 1986
Geosesarma johnsoni (Serène, 1968)
Geosesarma katibas Ng, 1995
Geosesarma krathing Ng & Naiyanetr, 1992
Geosesarma larsi Ng & Grinang, 2018
Geosesarma lawrencei Manuel-Santos & Yeo, 2007
Geosesarma leprosum (Schenkel, 1902)
Geosesarma maculatum (De Man, 1892)
Geosesarma malayanum Ng & Lim, 1986
Geosesarma nannophyes (De Man, 1885)
Geosesarma nemesis Ng, 1986
Geosesarma noduliferum (De Man, 1892)
Geosesarma notophorum Ng & C. G. S. Tan, 1995
Geosesarma ocypodum (Nobili, 1899)
Geosesarma penangense (Tweedie, 1940)
Geosesarma peraccae (Nobili, 1903)
Geosesarma protos Ng & Takeda, 1992
Geosesarma rathbunae (Serène, 1968)
Geosesarma rouxi (Serène, 1968)
Geosesarma sabanum Ng, 1992
Geosesarma sarawakense (Serène, 1968)
Geosesarma scandens Ng, 1986
Geosesarma serenei Ng, 1986
Geosesarma solomonense (Serène, 1968)
Geosesarma starmuhlneri Pretzmann, 1984
Geosesarma sumatraense Ng, 1986
Geosesarma sylvicola (De Man, 1892)
Geosesarma ternatense (Serène, 1968)
Geosesarma teschi Ng, 1986
Geosesarma thelxinoe (De Man, 1908)
Geosesarma tiomanicum Ng, 1986
Geosesarma vicentense (Rathbun, 1914)

As of March 2015, professor Peter Ng of National University of Singapore has named 20 Geosesarma species, and he "has another half a dozen or so newly collected Geosesarma species from Southeast Asia in his lab, and these species still need to be named and described."[4][5]
Threats

Geosesarma dennerle and Geosesarma hagen, both originally from Java, are threatened by illegal overcollection for the aquarium trade.[6]
References

Peter Davie (2012). "Geosesarma de Man, 1892". WoRMS. World Register of Marine Species. Retrieved November 8, 2012.
Hartnoll, Richard G. (1998). "Evolution, systematics, and geographical distribution". In Warren W. Burggren; Brian R. McMahon (eds.). Biology of the Land Crabs. Cambridge University Press. pp. 6–54. ISBN 9780521306904.
Pati, S. K.; Dev Roy, M. K.; Sharma, R. M. "Freshwater crabs" (PDF). Checklist of Indian fauna. Zoological Survey of India. Archived from the original (PDF) on September 17, 2013. Retrieved November 8, 2012.
Blaszczak-Boxe, Agata (March 19, 2015). "Mystery of the 'Vampire Crabs' Solved". livescience.com.
Blaszczak-Boxe, Agata (19 March 2015). "Mystery of the 'Vampire Crabs' Solved". LiveScience. Retrieved 2021-02-11.
Mahbu, Amri (March 23, 2015). "New Species of Javan Vampire Crabs Face Potential Exploitation".

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