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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Crustacea
Superclassis: Multicrustacea
Classis: Malacostraca
Subclassis: Eumalacostraca
Superordo: Eucarida
Ordo: Decapoda
Subordo: Pleocyemata
Infraordo: Brachyura
Sectio: Eubrachyura
Subsectio: Heterotremata
Superfamilia: Aethroidea

Familia: Aethridae
Genus: Hepatus
Species (15): H. chiliensis – H. epheliticus – H. gronovii – H. kossmanni – H. lineatus – H. pudibundus – H. scaber – †H. biformis – †H. bottomsi – †H. guraboensis – †H. lineatinus – †H. lineatus – †H. nodosus – †H. praecox – †H. spinimarginatus

[source: WoRMS, except for fossil species]

Hepatus Latreille, 1802

Primary references

Latreille, P.A. 1802: Histoire naturelle, générale et particulière, des Crustacés et des Insectes. Vol. 5: 1–407, pls. 38–43. Vol. 6: 1–392, pls. 42–57. Paris, Dufart.

Hepatus is a genus of crabs in the family Aethridae, containing seven extant species,[1] plus some fossil species:[2]

Hepatus chiliensis H. Milne-Edwards, 1837
Hepatus epheliticus (Linnaeus, 1763)
Hepatus gronovii Holthuis, 1959
Hepatus kossmanni Neumann, 1878
Hepatus lineatus Rathbun, 1898
Hepatus pudibundus (Herbst, 1785)
Hepatus scaber Holthuis, 1959

Both H. chiliensis and H. lineatus are also known as fossils.[2] The other fossil species include:[3]

Hepatus bottomsi Blow, 2003
Hepatus lineatus Rathbun, 1898
Hepatus nodosus Collins & Morris, 1976
Hepatus praecox Collins et al., 1996
Hepatus spinimarginatus Feldmann et al., 2005


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P. Davie (2010). "Hepatus Latreille, 1802". World Register of Marine Species. Retrieved December 6, 2010.
Sammy De Grave; N. Dean Pentcheff; Shane T. Ahyong; et al. (2009). "A classification of living and fossil genera of decapod crustaceans" (PDF). Raffles Bulletin of Zoology. Suppl. 21: 1–109. Archived from the original (PDF) on 2011-06-06.
Rodney M. Feldmann; Carrie E. Schweitzer; Alfonso Encinas (2005). "New decapods from the Navidad Formation (Miocene) of Chile" (PDF). Journal of Crustacean Biology. 25 (3): 427–449. doi:10.1651/c-2547.


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