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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Cladus: Macroheterocera
Superfamilia: Geometroidea

Familia: Geometridae
Subfamilia: Larentiinae
Tribus: Asthenini
Genera: AgnibesaAnydreliaAsthenaAsthenotrichaBihastinaChaetolophaEoisEpicymeEschatarchiaEuchoeca – Hastina – HydreliaLaciniodes – Leucoctenorrhoe – Macrohastina – Minoa – Nomenia – Palpoctenidia – Parachaetolopha – Parasthena – Poecilasthena – Polynesia – Pseudostegania – Venusia



Han, H.X. , D. Stüning & D.Y. Xue, 2010: Taxonomic review of the genus Pseudostegania Butler, 1881, with description of four new species and comments on its tribal placement in the Larentiinae (Lepidoptera: Geometridae). Entomological Science 13 (2): 234–249. DOI: 10.1111/j.1479-8298.2010.00379.x.
Schmidt, O., 2002: A revision of the genus Chaetolopha Warren (Lepidoptera : Geometridae : Larentiinae) with a description of Parachaetolopha, gen. nov. Invertebrate Systematics, 16 (5): 703–733. Abstract: DOI: 10.1071/IT01013
Viidalepp, J. 2011: A morphological review of tribes in Larentiinae (Lepidoptera: Geometridae). Zootaxa, 3136: 1–44. Preview
Xue, D.Y.; Scoble, M.J., 2002: A review of the genera associated with the tribe Asthenini (Lepidoptera: Geometridae: Larentiinae). Bulletin of the Natural History Museum (entomology series), 71 (1): 77–133. Abstract: DOI: 10.1017/S0968045402000044.

Asthenini is a tribe of geometer moths under subfamily Larentiinae first described by Warren in 1893. The tribe has been combined with Eupitheciini in the past, most notably by Jeremy Daniel Holloway in his work The Moths of Borneo.[1]


The tribe is represented in all the major biogeographical regions, but is most diverse in eastern Asia and is poorly represented in the Afrotropics. It is also poorly represented in the Neotropics if the genus Eois is excluded.

227 species group taxa (species and subspecies) were included in the tribe in 2002. Later research re-included two genera, including the large genus Eois (about 250 species), more than doubling the number of species.
Recognized genera and species

Critical analysis by Dayong Xue and Malcolm J. Scoble [2] indicated 18 genera that belong to the Asthenini with 1 additional, ungrouped species. The 18 genera recognized by this research are:

Agnibesa Moore 1888
Anydrelia Prout 1938
Asthena Hübner 1825
Asthenotricha Warren 1899
Bihastina Prout 1916
Epicyme Meyrick 1885
Eschatarchia Warren 1894
Euchoeca Hübner 1823
Hastina Moore 1888
Hydrelia Hübner 1825
Leucoctenorrhoe Warren 1904
Macrohastina Inoue 1982
Nomenia Pearsall 1905
Palpoctenidia Prout 1930
Parasthena Warren 1902
Poecilasthena Warren 1894
Polynesia Swinhoe 1892
Venusia Curtis 1839

Later research re-included the following genera:

Eois (but see below)
Minoa (but see below)
Pseudostegania Butler, 1881

Disassociated genera

The following 7 genera are sometimes associated with the Asthenini, but were excluded by the work of Xue and Scoble:


Eois and Minoa were excluded by Xue and Scoble, but later again included by Viidalepp in 2011.
Species incertae sedis

Species 'Chalyboclydon' flexilinea was identified by Xue and Scoble as being part of the Asthenini. However, it has not been associated with a specific Asthenini genus.

Holloway, Jeremy Daniel. "Part 10 Family Geometridae, Subfamilies Sterrhinae and Larentiinae". The Moths of Borneo. Retrieved March 13, 2019.
Xue, Dayong; Scoble, Malcolm J. (June 27, 2002). "A review of the genera associated with the tribe Asthenini (Lepidoptera: Geometridae: Larentiinae)". Bulletin of the Natural History Museum. Entomology Series. 71 (1): 77–133. doi:10.1017/S0968045402000044.

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