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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Cleroidea
Familiae (18 + 1†): AcanthocnemidaeBiphyllidaeByturidaeChaetosomatidaeCleridae – Lophocateridae – MauroniscidaeMelyridae – Peltidae – PhloiophilidaePhycosecidaePrionoceridae – Protopeltidae – Rentoniidae – Rhadalidae – Thanerocleridae – Thymalidae – Trogossitidae – †Boleopsidae
[source: Gimmel et al. (2019)]
Genera incertae sedis

†Forticatinius – †Mathesius


Cleroidea Latreille, 1802
Primary references

Latreille, P.A. 1802. Histoire naturelle, générale et particulière des crustacés et des insectes. Ouvrage faisant suite à l’histoire naturelle générale et particulière, composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, membre de plusieurs sociétés savantes. Familles naturelles des genres. Tome troisième. F. Dufart, Paris, xii + pp. 13–467 + [1 (errata)]. BHL Reference page.

Additional references

Beutel, R.G.; Pollock, D.A. 2000: Larval head morphology of Phycosecis litoralis (Pascoe) (Coleoptera: Phycosecidae) with phylogenetic implications. Invertebrate taxonomy, 14: 825–835. DOI: 10.1071/IT00030
Bocakova, M., Constantin, R. & Bocak, L. 2012 (online 2011). Molecular phylogenetics of the melyrid lineage (Coleoptera: Cleroidea). Cladistics 28(2): 117–129. DOI: 10.1111/j.1096-0031.2011.00368.x Open access. Reference page.
Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Chittaro, Y. & Sanchez, A. 2019. Liste commentée des Cleroidea (Coleoptera) de Suisse [Annotated checklist of Cleroidea (Coleoptera) of Switzerland]. Alpine Entomology 3: 141–167. DOI: 10.3897/alpento.3.35994 Reference page.
Crowson, R.A. 1964. A review of the classification of Cleroidea (Coleoptera), with descriptions of two new genera of Peltidae and of several new larval types. Transactions of the Royal Entomological Society of London 116(12): 275–327, plate 1. DOI: 10.1111/j.1365-2311.1964.tb02298.x Paywall. Reference page.
Crowson, R.A. 1970. Further observations on Cleroidea (Coleoptera). Proceedings of the Royal Entomological Society of London (B) 39(1–2): 1–20. DOI: 10.1111/j.1365-3113.1970.tb00247.x Reference page.
Gimmel, M.L., Bocakova, M., Gunter, N.L. & Leschen, R.A.B. 2019. Comprehensive phylogeny of the Cleroidea (Coleoptera: Cucujiformia). Systematic Entomology 44(3): 527–558. DOI: 10.1111/syen.12338 Paywall. Reference page.
Hájek, J. & Švihla, V. 2012. Catalogue of the type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic. Trogossitidae, Cleridae, Prionoceridae, Melyridae, Dasytidae, Rhadalidae and Malachiidae. Acta Entomologica Musei Nationalis Pragae 52(2): 603–654. PDF Reference page.
Kirejtshuk, A.G., Ponomarenko, A.G., Prokin, A.A., Chang, H., Nikolajev, G.V. & Ren, D. 2010. Current knowledge of Mesozoic Coleoptera from Daohugou and Liaoning (northeast China). Acta Geologica Sinica - English Edition 84(4): 783–792. DOI: 10.1111/j.1755-6724.2010.00253.x Paywall. Reference page.
Kolibáč, J. 2004. Metaxinidae fam. nov., a new family of Cleroidea (Coleoptera). Entomologica Basiliensia 26: 239–268. ResearchGate Open access. Reference page.
Kolibáč, J. & Huang, D.-Y. 2011. Mathesius liaoningensis gen. et sp. nov. of Jehol Biota, a presumptive relative of the clerid or thaneroclerid branches of Cleroidea (Coleoptera). Zootaxa 2872: 1–17. Preview DOI: 10.11646/zootaxa.2872.1.1 Paywall. Reference page.
Leschen, R.A.B. 2010: 9. Cleroidea Latreille, 1802. Introduction and phylogeny. Pp. 237-239 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755
I.Löbl & A.Smetana (eds). 2007 Catalogue of Palearctic Coleoptera. Vol. 4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea and Cucujoidea. Apollo Books, Stenstrup, Denmark ISBN 87-88757-67-6, p. 363
Mirutenko, V. & Ghahari, H. 2018. Contribution to study of the superfamily Cleroidea (Coleoptera) from Iran and adjacent countries: a checklist of Melyridae, Prionoceridae and Trogossitidae. Zootaxa 4378(1): 40–48. DOI: 10.11646/zootaxa.4378.1.2 Reference page.
Plonski, I.S. & Ghahari, H. 2018. An annotated checklist of the Iranian Dasytidae and Rhadalidae (Coleoptera: Cleroidea). Zootaxa 4472(3): 452–470. DOI: 10.11646/zootaxa.4472.3.2 Paywall Reference page.
Tan, J.J.; Ren, D. 2007: Two exceptionally well-preserved catiniids (Coleoptera: Archostemata: Catiniidae) from the late mesozoic of northeastern China. Annals of the Entomological Society of America, 100: 666–672. DOI: 10.1603/0013-8746(2007)100[666:TEWCCA]2.0.CO;2 PDF


Australian Faunal Directory
Tree of Life Web Project. 1995. Cleroidea. Version 01 January 1995 (temporary) in The Tree of Life Web Project
Atlas of beetles of Russia: the superfamily CLEROIDEA

Vernacular names
suomi: Kirjokuoriasmaiset
magyar: Szúfarkasszerűek
日本語: カッコウムシ上科
中文: 郭公蟲總科

Cleroidea is a small superfamily of beetles containing about 10,000 species. Most of the members of the group are somewhat slender, often with fairly soft, flexible elytra, and typically hairy or scaly.

Families included are:[1]

Acanthocnemidae Crowson, 1964
Biphyllidae LeConte, 1861
†Boleopsidae Kirejtshuk & Nel, 2013
Byturidae Gistel, 1848
Chaetosomatidae Crowson, 1952 (incl. Metaxinidae)
Cleridae Latreille, 1802 (checkered beetles)
Lophocateridae Crowson, 1964
Mauroniscidae Majer, 1995
Melyridae Leach, 1815 (soft-winged flower beetles) (incl. Attalomimidae, Malachiidae and Dasytidae)
Peltidae Latreille, 1806
Phloiophilidae Kiesenwetter, 1863
Phycosecidae Crowson, 1952
Prionoceridae Lacordaire, 1857
Protopeltidae Crowson, 1966
Rentoniidae Crowson, 1966
Rhadalidae LeConte, 1861 (incl. Gietellidae)
Thanerocleridae Chapin, 1924
Thymalidae Léveillé, 1888
Trogossitidae Latreille, 1802 (bark-gnawing beetles)

Most species belong to the families Cleridae and Melyridae, followed by Trogossitidae.

Gimmel, Matthew L.; Bocakova, Milada; Gunter, Nicole L.; Leschen, Richard A.B. (2019). "Comprehensive phylogeny of the Cleroidea (Coleoptera: Cucujiformia)". Systematic Entomology. 44 (3): 527–558. doi:10.1111/syen.12338.

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