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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Cleroidea

Familia: Biphyllidae
Genera (7): AlthaesiaAnchoriusAnobocaelusBiphyllusDiplocoelusEuderopusGonicoelus


Biphyllidae LeConte, 1861

Diphyllidae LeConte, 1861: 105

Primary references

LeConte, J.L. 1861. Classification of the Coleoptera of North America. Part I. Prepared for the Smithsonian Institution. Smithsonian Miscellaneous Collections 3: 1–208. [pp. 209–286 published in March 1862] BHL Reference page.

Additional references

Cline, A.R.; McHugh, J.V. 2010: New generic synonymy in Biphyllidae (Coleoptera: Cucujoidea), with a checklist of Anchorius Casey, 1900. Coleopterists bulletin, 64: 98–99. DOI: 10.1649/0010-065X-64.1.98
Cline, A.R.; Shockley, F.W. 2010: 10.6. Biphyllidae LeConte, 1861. Pp. 306-311 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755
Goodrich, M.A.; Springer, C.A. 1992: A revision of the family Biphyllidae (Coleoptera) for America, north of Mexico. Coleopterists bulletin, 46: 361–377. JSTOR
I.Löbl & A.Smetana (eds). 2007 Catalogue of Palearctic Coleoptera. Vol. 4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea and Cucujoidea. Apollo Books, Stenstrup, Denmark ISBN 87-88757-67-6, p. 547
Vitali, F. 2010: Diplocoelus probiphyllus n. sp., the first known fossil false skin beetle (Coleoptera: Biphyllidae). Annales de la Société entomologique de France (n.s.), 46: 168–172. ISSN: 0037-9271


Australian Faunal Directory (stub)
Atlas of false skin beetles (Biphyllidae) of Russia

Vernacular names
English: False Skin Beetles

Biphyllidae, or false skin beetles, are a family of beetles, in the suborder Polyphaga. Worldwide, about 195 species are known. They live under the bark of dead trees and feed on fungi.[1] It contains the following genera:[2]

Althaesia Pascoe, 1860
Anchorius Casey, 1900
Anobocaelus Sharp, 1902
Biphyllus Dejean, 1821
Diplocoelus Guérin-Méneville, 1836
Euderopus Sharp, 1900
Gonicoelus Sharp, 1900
†Paleobiphyllus Makarov and Perkovsky 2019[3] Taimyr amber, Russia, Santonian


Michael A. Ivie (2002). Ross H. Arnett & Michael Charles Thomas (ed.). American Beetles: Polyphaga: Scarabaeoidea through Curculionoidea. Volume 2 of American Beetles. CRC Press. ISBN 978-0-8493-0954-0.
Biphyllidae Species List at Joel Hallan’s Biology Catalog. Texas A&M University. Retrieved on 15 May 2012.
Makarov, Kirill V.; Perkovsky, Evgeny E. (February 2020). "Smallest and oldest false skin beetle: Paleobiphyllus ponomarenkoi gen. et sp. nov. (Coleoptera: Cleroidea: Biphyllidae) from Santonian Taimyr amber, northern Russia". Cretaceous Research. 106: 104238. doi:10.1016/j.cretres.2019.104238.

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