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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea
Familiae (12 + 5†): BelohinidaeDiphyllostomatidae – Eremazidae – GeotrupidaeGlaphyridaeGlaresidaeHybosoridaeLucanidaeOchodaeidaePassalidaePleocomidaeScarabaeidaeTrogidae – †Alloioscarabaeidae – †Lithoscarabaeidae – †Paralucanidae – †Passalopalpidae – †Septiventeridae
[sources: Smith (2006), Krell (2007), Bai et al. (2012a, b)]

Genera incertae sedis (7): †Aliscarabaeus – †Avitortor – †Cretoserica – †Hongscarabaeus – †Hybosorites – †Mesoscarabaeus – †Opiselleipon

Scarabaeoidea Latreille, 1802


Ahrens, D., Schwarzer, J. & Vogler, A.P. 2014. The evolution of scarab beetles tracks the sequential rise of angiosperms and mammals. Proceedings of the Royal Society (B), 281(1791) DOI: 10.1098/rspb.2014.1470 Reference page.
Bai, M. et al. 2012 (Online) 2013 (Print): Septiventeridae, a new and ancestral fossil family of Scarabaeoidea (Insecta: Coleoptera) from the Late Jurassic to Early Cretaceous Yixian Formation. Journal of systematic palaeontology, 11(3): 359–374. DOI: 10.1080/14772019.2012.660995 ZooBank
Bai, M., Ahrens, D., Yang, X-K. & Ren, D. 2012. New fossil evidence of the early diversification of scarabs: Alloioscarabaeus cheni (Coleoptera: Scarabaeoidea) from the Middle Jurassic of Inner Mongolia, China. Insect science 19(2): 159–171. DOI: 10.1111/j.1744-7917.2011.01460.x Open access. Reference page.
Bezděk, A. & Hájek, J. 2009. Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic - Scarabaeoidea: Bolboceratidae, Geotrupidae, Glaphyridae, Hybosoridae, Ochodaeidae and Trogidae. Acta Entomologica Musei Nationalis Pragae 49(1): 297–332. PDF Reference page.
Browne, D.J.; Scholtz, C.H. 1995: Phylogeny of the families of Scarabaeoidea (Coleoptera) based on characters of the hindwing articulation, hindwing base and wing venation. Systematic entomology, 20: 145–173. DOI: 10.1111/j.1365-3113.1995.tb00089.x
Browne, J.; Scholtz, C.H. 1999: A phylogeny of the families of Scarabaeoidea (Coleoptera). Systematic entomology, 24: 51–84. DOI: 10.1046/j.1365-3113.1999.00067.x
Byk, A., Matusiak, A, Taszakowski, A, Szczepański, W.T., Walczak, M., Bunalski, M. & Karpiński, L. 2020. New and interesting findings of scarab beetles (Coleoptera, Scarabaeoidea) from Tajikistan. ZooKeys, 1003: 57–82. DOI: 10.3897/zookeys.1003.55457 Open access Reference page.
Cosandey, V., Sanchez, A. & Chittaro, Y. 2017. Liste commentée des Scarabaeoidea (Coleoptera) de Suisse. Alpine Entomology 1: 57–90. [In French] DOI: 10.3897/alpineentomology.1.21179. Reference page.
Grebennikov, V.V.; Scholtz, C.H. 2004: The basal phylogeny of Scarabaeoidea (Insecta: Coleoptera) inferred from larval morphology. Invertebrate systematics, 18: 321–348.
Kaneko, N. & Wada, K. 2020. Catalogue of the type material of Scarabaeoidea (Coleoptera) deposited in the Research Institute of Evolutionary Biology, Tokyo, Japan. ZooKeys, 958: 35–89. DOI: 10.3897/zookeys.958.52799 Open access Reference page.
Krajcik, M. 2012: Checklist of the World Scarabaeoidea. Animma.X Supplement, (5) [not seen]
Krell, F.-T. 2006: Fossil record and evolution of Scarabaeoidea (Coleoptera: Polyphaga). Coleopterists bulletin, 60(sp5): 120–143. DOI: 10.1649/0010-065X(2006)60[120:FRAEOS]2.0.CO;2[broken/unregistered!] PDF
Krell, F.-T. 2007: Catalogue of fossil Scarabaeoidea (Coleoptera: Polyphaga) of the Mesozoic and Tertiary – Version 2007 –. Denver Museum of Nature and Science technical report, 2007-8: 1–79. PDF
Latreille, P.A. 1802: Histoire naturelle, générale et particulière des Crustacés et des Insectes. Tome 3. F. Dufart, Paris. Internet Archive BHL
Scholtz, C.H.; Grebennikov, V.V. 2005: 12. Scarabaeiformia/13. Scarabaeoidea. Pp. 345-425 in: Beutel, R.G. & Leschen, R.A.B. (volume eds.) Coleoptera, Beetles. Volume 1: Morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Smith, A.B.T. 2006: A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society monograph, 5: 144–204. DOI: 10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2 PDF
Smith, A.B.T.; Hawks, D.C.; Heraty, J.M. 2006: An overview of the classification and evolution of the major scarab beetle clades (Coleoptera: Scarabaeoidea) based on preliminary molecular analyses. Coleopterists Society monograph, 5: 35–46.
Catalogue of Palearctic Coleoptera. Vol. 3 ed. I. Lobl, & A. Smetana, Apollo Books, Stenstrup, Denmark, 2006
ISBN 87-88757-59-5, p.63


Browne, D. Jonathan and Clarke H. Scholtz 1995. Scarabaeiformia. Scarabaeoidea. Scarabs, stag beetles, dung beetles, rain beetles, etc. Version 01 January 1995 (under construction) in The Tree of Life Web Project
Atlas of beetles of Russia: the superfamily SCARABAEOIDEA
Australian Faunal Directory

Vernacular names
日本語: コガネムシ上科
한국어: 풍뎅이상과
русский: Скарабеоидные
中文: 金龜子總科

Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year.[1] Its constituent families are also undergoing revision presently, and the family list below is only preliminary.


The following families are listed in Bouchard (2011):[2]

Belohinidae Paulian, 1959
Diphyllostomatidae Holloway, 1972 (false stag beetles)
Geotrupidae Latreille, 1802 (earth-boring dung beetles)
Glaphyridae MacLeay, 1819 (bumble bee scarab beetles)
Glaresidae Kolbe, 1905 (enigmatic scarab beetles)
Hybosoridae Erichson, 1847 (scavenging scarab beetles)
inclusive of Ceratocanthidae White, 1842 (pill scarab beetles)
Lucanidae Latreille 1804 (stag beetles)
Ochodaeidae Mulsant and Rey 1871 (sand-loving scarab beetles)
Passalidae Leach, 1815 (bess beetles)
Pleocomidae LeConte 1861 (rain beetles)
Scarabaeidae Latreille 1802 (scarab beetles)
Trogidae MacLeay 1819 (hide beetles)
† Coprinisphaeridae Genise, 2004
† Pallichnidae Genise, 2004
† Passalopalpidae Boucher, et al., 2016

See also

List of subgroups of the order Coleoptera

In Peru alone, there are 1042 known species of Scarabaeoidea as of 2015.[3] This is due to Peru's high biodiversity and endemism.

Brett C. Ratcliffe (2002). "A checklist of the Scarabaeoidea (Coleoptera) of Panama" (PDF). Zootaxa (32): 1–48.
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys. Pensoft Publishers (88): 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
Ratcliffe, B. C.; Jameson, M. L.; Figueroa, L.; Cave, R. D.; Paulsen, M. J.; Cano, Enio B.; Beza-Beza, C.; Jimenez-Ferbans, L.; Reyes-Castillo, P. (2015). "Beetles (Coleoptera) of Peru: A Survey of the Families. Scarabaeoidea". Journal of the Kansas Entomological Society. 88 (2): 186–207. doi:10.2317/kent-88-02-186-207.1.

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