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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamiliae: Scarabaeoidea

Overview of familiae

BelohinidaeDiphyllostomatidaeGeotrupidaeGlaphyridaeGlaresidaeHybosoridaeLucanidaeOchodaeidaePassalidaePleocomidaeScarabaeidaeTrogidae – †Lithoscarabaeidae – †Paralucanidae

[list of familiae after Smith (2006), Krell (2007)]

Caterino, M.S.; Hunt, T.; Vogler, A.P. 2005: On the constitution and phylogeny of Staphyliniformia (Insecta: Coleoptera). Molecular phylogenetics and evolution, 34: 655–672.
Gerlach, J. 2009: Series Scarabaeiformia. Pp. 79-86 in: Gerlach, J. (ed.) The Coleoptera of the Seychelles Islands. Pensoft Publishers, Sofia-Moscow. ISBN 9789546424983
Krell, F.-T. 2007: Catalogue of fossil Scarabaeoidea (Coleoptera: Polyphaga) of the Mesozoic and Tertiary – Version 2007 –. Denver Museum of Nature and Science technical report, 2007-8: 1–79. PDF
McKenna, D.D. et al. 2014 (Online) 2015 (Print): Phylogeny and evolution of Staphyliniformia and Scarabaeiformia: forest litter as a stepping stone for diversification of nonphytophagous beetles. Systematic entomology 40(1): 35–60. DOI: 10.1111/syen.12093 Reference page.
Scholtz, C.H.; Grebennikov, V.V. 2005: 12. Scarabaeiformia/13. Scarabaeoidea. Pp. 345-425 in: Beutel, R.G. & Leschen, R.A.B. (volume eds.) Coleoptera, Beetles. Volume 1: Morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Smith, A.B.T. 2006: A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society monograph, 5: 144–204.PDF


Browne, D. Jonathan and Clarke H. Scholtz 1995. Scarabaeiformia. Scarabaeoidea. Scarabs, stag beetles, dung beetles, rain beetles, etc. Version 01 January 1995 (under construction) in The Tree of Life Web Project
Atlas of beetles of Russia: the superfamily SCARABAEOIDEA

Vernacular names
日本語: コガネムシ下目

Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year.[1] Its constituent families are also undergoing revision presently, and the family list below is only preliminary.


The following families are listed in Bouchard (2011):[2]

Belohinidae Paulian, 1959
Diphyllostomatidae Holloway, 1972 (false stag beetles)
Geotrupidae Latreille, 1802 (earth-boring dung beetles)
Glaphyridae MacLeay, 1819 (bumble bee scarab beetles)
Glaresidae Kolbe, 1905 (enigmatic scarab beetles)
Hybosoridae Erichson, 1847 (scavenging scarab beetles)
inclusive of Ceratocanthidae White, 1842 (pill scarab beetles)
Lucanidae Latreille 1804 (stag beetles)
Ochodaeidae Mulsant and Rey 1871 (sand-loving scarab beetles)
Passalidae Leach, 1815 (bess beetles)
Pleocomidae LeConte 1861 (rain beetles)
Scarabaeidae Latreille 1802 (scarab beetles)
Trogidae MacLeay 1819 (hide beetles)
† Coprinisphaeridae Genise, 2004
† Pallichnidae Genise, 2004
† Passalopalpidae Boucher, et al., 2016

See also

List of subgroups of the order Coleoptera

In Peru alone, there are 1042 known species of Scarabaeoidea as of 2015.[3] This is due to Peru's high biodiversity and endemism.

Brett C. Ratcliffe (2002). "A checklist of the Scarabaeoidea (Coleoptera) of Panama" (PDF). Zootaxa (32): 1–48.
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys. Pensoft Publishers (88): 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
Ratcliffe, B. C.; Jameson, M. L.; Figueroa, L.; Cave, R. D.; Paulsen, M. J.; Cano, Enio B.; Beza-Beza, C.; Jimenez-Ferbans, L.; Reyes-Castillo, P. (2015). "Beetles (Coleoptera) of Peru: A Survey of the Families. Scarabaeoidea". Journal of the Kansas Entomological Society. 88 (2): 186–207. doi:10.2317/kent-88-02-186-207.1.

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