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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Orthoptera
Subordo: Ensifera
Infraordo: Tettigoniidea
Superfamilia: Tettigonioidea

Subfamilia: Conocephalinae
Tribus: Copiphorini
Genera: Acanthacara - Acantheremus - Apoecides - Artiotonus – Banza - Belocephalus - Borinquenula - Bucrates - Caetitus – Caulopsis - Clasma - Conocephaloides - Copiphora - Coryphodes - Daedalellus - Dectinomima - Dorycoryphus - Erioloides - Eriolus - Eucaulopsis - Euconocephalus - Eurymetopa - Gryporhynchium - Lamniceps - Lanista - Liliella - Liostethus - Lirometopum - Loboscelis - Mayacephalus – Melanophoxus - Metacaputus - Moncheca - Montesa - Mygalopsis - Neoconocephalus - Ocana - Oxyprora - Panacanthus - Parabucrates - Paroxyprora - Pedinostethus - Phaneracra - Phoxacris - Plastocorypha - Pluviasilva - Poascirtus - Podacanthophorus - Pseudorhynchus - Pyrgocorypha - Ruspolia - Sphodrophoxus - Toledopizia - Vestria - Xestophrys

Copiphorini Karny, 1912

Copiphorinae Karny, 1912


Karny, H. 1912. Gallenbewohnende Thysanopteren aus Java. Marcellia 11: 115–169. BHL Reference page.
Antunes, A.F., Takiya, D.M. & Chamorro-Rengifo, J. 2018. Caetitus gen. nov.: a new genus of Neotropical Copiphorini (Tettigoniidae: Conocephalinae: Copiphorini) to include C. porteri (Bolivar, 1903). Zootaxa 4375(4): 578–586. DOI: 10.11646/zootaxa.4375.4.7 Reference page.
Cadena-Castañeda, O.J., Sierra, J.M. & Cortés-Torres, C. 2016. Studies in Guatemalan Ensifera: Mayacephalus (Orthoptera: Tettigoniidae) a new cone-headed katydid genus. Zootaxa 4084(2): 293–300. DOI: 10.11646/zootaxa.4084.2.9.Reference page.
Chamorro-Rengifo, J.; Braun, H. 2010: The Tettigoniidae (Orthoptera) described by Salvador de Toledo Piza Jr. and deposited in the collection of the University of São Paulo, Escola Superior de Agricultura "Luiz de Queiroz", Brazil. Zootaxa, 2635: 41–66. Preview
Fontana, P.; Mariño-Pérez, R.; Woller, D.A. 2013: Studies in Mexican Tettigoniidae: A new genus of Copiphorini and the first description of male Conocephalus (Aphauropus) leptopterus Rehn and Hebard and the female of Insara acutitegmina Fontana, Buzzetti, Mariño-Pérez & García García. Zootaxa 3737(4): 429–453. DOI: 10.11646/zootaxa.3737.4.6 Reference page.
Liu, X.-W. et al. 2012. A study on the genus Pyrgocorypha Stål with description of one new species from China (Orthoptera, Tettigonoidea, Conocephalidae). Acta zootaxonomica sinica 37(1): 111–118. Reference page.


Copiphorini – Taxon details on National Center for Biotechnology Information (NCBI).
Orthoptera Species File Online : tribe Copiphorini Karny, 1912

The Copiphorini are a tribe of bush crickets or katydids in the family Tettigoniidae. Previously considered a subfamily (the Copiphorinae), they are now placed in the subfamily Conocephalinae.[1] Like some other members of Conocephalinae, they are known as coneheads, grasshopper-like insects with an extended, cone-shaped projection on their heads that juts forward in front of the base of the antennae.[2]

Species in the Copiphorini vary in length from about 24 to 74 mm (1 to 3 in). In most species, the female is considerably larger than the male, and in some, the largest male is smaller than the smallest female.[2] Like other members of the cricket suborder Ensifera, they differ from grasshoppers (suborder Caelifera) in having filamentous antennae that are longer than their bodies. Most species have loud songs which enable them to be identified. The call is made in most instances by the two fore wings rubbing together. Some species can hear with the aid of hearing organs in the tibia segments of their legs. The ovipositor is straight and slender and bears no teeth.[3]

Genera include:[4]

Acantheremus Karny, 1907
Anacaona Yong, 2019
Apoecides Bolívar, 1914
Artiotonus Montealegre-Z., Morris, Sarria-S. & Mason, 2011
Banza Walker, 1870
Belocephalus Scudder, 1875
Borinquenula Walker & Gurney, 1972
Brachycaulopsis Fontana, Mariño-Pérez & Woller, 2013
Bucrates Burmeister, 1838
Caetitus Antunes, Chamorro-Rengifo & Takiya, 2018
Caulopsis Redtenbacher, 1891
Clasma Karsch, 1893
Conocephaloides Perkins, 1899
Copiphora Serville, 1831
Coryphodes Redtenbacher, 1891
Daedalellus Uvarov, 1940
Dorycoryphus Redtenbacher, 1891
Erioloides Hebard, 1927
Eriolus Bolívar, 1888
Eucaulopsis Hebard, 1931
Euconocephalus Karny, 1907
Eurymetopa Redtenbacher, 1891
Gryporhynchium Uvarov, 1940
Lamniceps Bolívar, 1903
Lanista Bolívar, 1890
Lanistoides Sjöstedt, 1913
Liostethus Redtenbacher, 1891
Lirometopum Scudder, 1875
Loboscelis Redtenbacher, 1891
Mayacephalus Cadena-Castañeda, Monzón-Sierra & Cortés-Torres, 2016
Melanophoxus Karny, 1907
Metacaputus Naskrecki, 2000
Moncheca Walker, 1869
Montesa Walker, 1869
Mygalopsis Redtenbacher, 1891
Neoconocephalus Karny, 1907
Oxyprora Stål, 1873
Panacanthus Walker, 1869
Parabucrates Scudder, 1897
Paroxyprora Karny, 1907
Parvarhynchus Farooqi & Usmani, 2020
Pedinostethus Redtenbacher, 1891
Phaneracra Uvarov, 1936
Phoxacris Karny, 1907
Plastocorypha Karsch, 1896
Pluviasilva Naskrecki, 2000
Poascirtus Saussure, 1899
Podacanthophorus Naskrecki, 2000
Pseudorhynchus Serville, 1838
Pyrgocorypha Stål, 1873
Rombophora Yong, 2019
Ruspolia Schulthess, 1898
Santandera Koçak & Kemal, 2008
Sphodrophoxus Hebard, 1924
Toledopizia Chamorro-Rengifo & Braun, 2010
Unicorniella Yong, 2019
Vestria Stål, 1874
Xestophrys Redtenbacher, 1891


Orthoptera species file (retrieved 3 January 2018)
Walker, Thomas J. "Subfamily Copiphorinae". Singing Insects of North America. Retrieved 2014-07-24.
Walker, Thomas J. "How to recognize crickets, katydids, and cicadas". Singing Insects of North America. Retrieved 2014-07-24.
"subfamily Conocephalinae Burmeister, 1838". OSF Online. Retrieved 26 April 2022.

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