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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Orphninae
Tribus (2): Aegidiini - Orphnini
Overview of genera (17)

Aegidiellus – Aegidinus – Aegidium – Chaetonyx – Craniorphnus – Goniorphnus – Hybaloides – Hybalus – Madecorphnus – Orphnus – Paraegidium – Premicrodispus – Pseudorphnus – Renorphnus – Scarabadania – Stenosternus – Triodontus

[source: Frolov (2012)]

Colby, J. 2009: Monographic revision of the genus Aegidinus Arrow (1904) and generic phylogeny of the world Orphninae (Coleoptera: Scarabaeidae: Orphninae). Insecta Mundi, 0076: 1–41. PDF
Frolov, A.V. 2008: A new apterous species of the genus Orphnus MacLeay (Coleoptera: Scarabaeidae: Orphninae) from South Africa. Zootaxa, 1855: 65–68. Abstract & excerpt
Frolov, A.V. 2009: New scarab-beetle species (Coleoptera, Scarabaeidae, Aphodiinae, Orphninae) from central Asia and southern Africa. Entomological review, 89: 685–688. DOI: 10.1134/S0013873809060062
Frolov, A.V. 2010: Revision of the Madagascan genus Madecorphnus Paulian (Coleoptera: Scarabaeidae: Orphninae). Journal of natural history, 44: 1095–1111. DOI: 10.1080/00222931003632765
Frolov, A.V. 2010: [A new species of the genus Triodontus (Coleoptera, Scarabaeidae, Orphninae) from Southern Madagascar.] Entomologicheskoe obozrenie, 89 (4): 835–839. [not seen, in Russian, English translation in Entomological review, 90 (8): 1031–1034. (2010) DOI: 10.1134/S0013873810080087]
Frolov, A.V. 2012: Diagnosis, classification, and phylogenetic relationships of the orphnine scarab beetles (Coleoptera, Scarabaeidae: Orphninae). Entomologicheskoe obozrenie, 91(2): 332–351. [in Russian, English translation in Entomological review, 92(7): 782–797. (2012) DOI: 10.1134/S0013873812070056] Reference page.
Frolov, A.V. 2013: Little known Orphninae (Coleoptera, Scarabaeidae) collected by the Madagascan expeditions of the California Academy of Sciences. Biodiversity data journal, 1: e1016. DOI: 10.3897/BDJ.1.e1016 Reference page.
Frolov, A.V.; Montreuil O. 2009: A new genus of Orphninae (Coleoptera: Scarabaeidae) from Madagascar. Zoosystematica Rossica, 18: 65–69.
Khaustov, A.A. & Frolov, A.V. 2018. A new species, new genus and new records of heterostigmatic mites (Acari: Heterostigmata) phoretic on scarab beetles of the subfamily Orphninae (Coleoptera: Scarabaeidae). Zootaxa 4514(2): 181–201. DOI: 10.11646/zootaxa.4514.2.3 Paywall Reference page.

Orphninae is a subfamily of beetles in the scarab beetle family, Scarabaeidae. There are two tribes in the family, the New World Aegidiini and the Old World Orphnini.[1] They are mostly tropical beetles.[2]

As of 2013 there are about 195 species in the subfamily. They are classified in 15 genera: 4 in the tribe Aegidiini and 11 in the Orphnini. In addition, the genus Stenosternus is provisionally placed in the Aegidiini.[2]

Taxa include:[1][3]

Tribe Aegidiini
Genus Aegidiellus
Genus Aegidium
Genus Aegidinus
Genus Paraegidium
Genus Stenosternus[2]
Tribe Orphnini
Genus Madecorphnus
Genus Pseudorphnus
Genus Renorphnus
Genus Triodontus


Jameson, M. L. and F. C. Ocampo. Orphninae. Generic Guide to New World Scarab Beetles. University of Nebraska State Museum. 2005 Version.
Frolov, A. (2013). Stenosternus Karsch, a possible link between Neotropical and Afrotropical Orphninae (Coleoptera, Scarabaeidae). ZooKeys 335 33-46.

Frolov, A. V. and O. Montreuil. (2009). A new genus of Orphninae (Coleoptera: Scarabaeidae) from Madagascar. Zoosystematica Rossica 18(1) 65-69.

External links

Key to New World Orphninae Genera. Generic Guide to New World Scarab Beetles. University of Nebraska State Museum.

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