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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Pieridae
Subfamilia: Pierinae
Tribus (6): AnthochariniElodininiLeptosiainiNepheroniiniPieriniTeracolini


Pierinae Swainson, 1820.

Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol.5A, Pt.4A. Assn. for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439pp. Reference page.
Liseki, S.D.; Vane-Wright, R.I. 2014: Butterflies (Lepidoptera: Papilionoidea) of Mount Kilimanjaro: family Pieridae, subfamily Pierinae. Journal of natural history, DOI: 10.1080/00222933.2014.886343 Reference page.
van der Poorten, G.; van der Poorten, N. 2014: New and revised descriptions of the immature stages of some butterflies in Sri Lanka and their larval food plants (Lepidoptera: Pieridae). Part 2: Subfamily Pierinae (in part). Tijdschrift voor entomologie 157(1): 1-25. DOI: 10.1163/22119434-00002036 Reference page.
Wahlberg, N., Rota, J., Braby, M.F., Pierce, N.E. & Wheat, C.W. 2014. Revised systematics and higher classification of pierid butterflies (Lepidoptera: Pieridae) based on molecular data. Zoologica scripta 43(6): 641–650. DOI: 10.1111/zsc.12075 Paywall. Reference page.
Williams, M.C., 2008: Checklist of Afrotropical Papilionoidea and Hesperoidea.

Vernacular names
日本語: モンシロチョウ亜科, シロチョウ亜科
中文: 粉蝶亞科

The Pierinae are a large subfamily of pierid butterflies. The subfamily is one of several clades of butterflies often referred to as the whites.


It includes the following species (additional species can be found under the tribes listed in the adjacent box):

Anthocharis belia – Morocco orange tip
Anthocharis cethura – desert orangetip
Anthocharis cethura catalina – Catalina orangetip
Anthocharis damone – eastern orange tip
Anthocharis euphenoides – Provence orange tip
Anthocharis gruneri – Grüner's orange tip
Anthocharis julia – southern Rocky Mountain orangetip
Anthocharis lanceolata – gray marble
Anthocharis limonea – Mexican orangetip
Anthocharis midea – falcate orangetip
Anthocharis sara – Sara's orangetip
Anthocharis sara thoosa – southwestern orangetip
Anthocharis scolymus – yellow tip
Anthocharis stella – Stella orangetip
Anthocharis stella browningi – Utah Stella orangetip
Euchloe belemia – green-striped white
Euchloe charlonia – greenish black-tip
Euchloe crameri – western dappled white
Euchloe olympia – Olympia marble
Phoebis avellaneda – red-splashed sulphur
Pieris angelika – Arctic white
Pieris brassicae – large white
Pieris marginalis – margined white
Pieris marginalis reicheli – Reichel's margined white
Pieris oleracea – mustard white
Pieris oleracea frigida – Newfoundland white
Pieris rapae – small white
Pieris virginiensis – West Virginia white butterfly


Wahlberg, N.; Rota, J.; Braby, M.F.; Pierce, N.E.; Wheat, C.W. (2014). "Revised systematics and higher classification of pierid butterflies (Lepidoptera: Pieridae) based on molecular data". Zoologica Scripta. 43 (6): 641–650. doi:10.1111/zsc.12075.

Further reading

Glassberg, Jeffrey (2001). Butterflies through Binoculars, The West
Guppy, Crispin S. and Shepard, Jon H. (2001). Butterflies of British Columbia
James, David G. and Nunnallee, David (2011). Life Histories of Cascadia Butterflies
Pelham, Jonathan (2008). Catalogue of the Butterflies of the United States and Canada
Pyle, Robert Michael (2002). The Butterflies of Cascadia

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