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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Tenebrionoidea

Familia: Pythidae
Genera: AnaplopusIschyomiusPriognathusPythoSphalma – †Pythoceropsis – ?Osphyoplesius – ?Trimitomerus


Pythidae Solier, 1834

I.Löbl & A.Smetana (eds). 2008 Catalogue of Palearctic Coleoptera. Vol. 5: Tenebrionoidea. Apollo Books, Stenstrup, Denmark
ISBN 87-88757-84-6, p. 414
Pollock, D.A. 2002: 114. PYTHIDAE Solier 1834. Pp. 537-539 in: Arnett, R.H., jr.; Thomas, M.C.; Skelley, P.E.; Frank, J.H. (eds.): American beetles. Volume 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press LLC, Boca Raton, Florida. limited preview
Pollock, D.A. 2010: 11.23. Pythidae Solier, 1834. Pp. 708-715 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755
Young, D.K. 1976: The systematic position of Sphalma quadricollis Horn (Coleoptera: Salpingidae: Pythini) as clarified by discovery of its larva. Coleopterists bulletin, 30: 227–232. JSTOR
Young, D.K. 1991: Pythidae (Tenebrionoidea). Pp. 539-541 in: Lawrence, J.F. (coordinator) Order Coleoptera, in: Stehr, F.W. (ed.) Immature insects, 2 . Kendall/Hunt Publishing Company, Iowa, USA.


Atlas of pythid beetles (Pythidae) of Russia

Vernacular names
Deutsch: Drachenkäfer
English: Pythid beetles / Dead log bark beetles
русский: Трухляки

The family Pythidae is a small group of beetles with no vernacular common name, though recent authors have coined the name dead log bark beetles. There are about 10 genera and 17 described species in Pythidae.[1][2][3][4]
Pytho kolwensis


These 10 genera belong to the family Pythidae:

Conomorphus Champion, 1889 g
Ischyomius Chevrolat, 1878 g
Osphyoplesius Winkler, 1915 g
Physiomorphus Pic, 1917 g
Priognathus LeConte, 1850 i c g b
Pytho Latreille, 1796 i c g b
Pythoceropsis Wickham, 1913 g
Sphalma Horn, 1872 i c g b
Techmessa Bates, 1874 g
Trimitomerus Horn, 1888 i c g b

Data sources: i = ITIS,[1] c = Catalogue of Life,[2] g = GBIF,[3] b =[4]

"Pythidae Report". Integrated Taxonomic Information System. Retrieved 2018-05-06.
"Browse Pythidae". Catalogue of Life. Retrieved 2018-05-06.
"Pythidae". GBIF. Retrieved 2018-05-06.

"Pythidae Family Information". Retrieved 2018-05-06.

Gustafsson, B. Catalogus Coleopterorum Sueciae (2004), Naturhistoriska riksmuseet, Stockholm.

Further reading

Arnett, R.H. Jr.; Thomas, M. C.; Skelley, P. E.; Frank, J. H., eds. (2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. ISBN 978-0849309540.
Bouchard, P. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 88: 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
Crotch, G.R. (1873). Check list of the Coleoptera of America, north of Mexico. Naturalists' Agency. doi:10.5962/bhl.title.38811. ISBN 0665070772.
Kubisz, Daniel; Iwan, Dariusz; Tykarski, Piotr (2014). Tenebrionoidea: Tetratomidae, Melandryidae, Ripiphoridae, Prostomidae, Oedemeridae, Mycteridae, Pythidae, Aderidae, Scraptiidae, Critical checklist, distribution in Poland and meta-analysis. University of Warsaw - Faculty of Biology & Natura optima dux Foundation. ISBN 978-83-935107-9-5.
Lawrence, J.F.; Newton Jr., A.F. (1995). Pakaluk, James; Slipinski, Stanislaw Adam (eds.). Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Vol. 2. Muzeum i Instytut Zoologii PAN. pp. 779–1006. ISBN 83-85192-34-4.
LeConte, J.L. (1861). Classification of the Coleoptera of North America. Smithsonian Miscellaneous Collections. Vol. 3. Smithsonian Institution. doi:10.5962/bhl.title.38459. ISBN 0665100558.
Misof, B.; Liu, S.; Meusemann, K.; Peters, R.S.; et al. (2014). "Phylogenomics resolves the timing and pattern of insect evolution". Science. 346 (6210): 763–7. Bibcode:2014Sci...346..763M. doi:10.1126/science.1257570. PMID 25378627. S2CID 36008925.
Pollock, Darren A. (2002). Arnett, Ross H. Jr.; Thomas, Michael C.; Skelley, Paul E.; Frank, J.H. (eds.). Family 114. Pythidae Solier 1834. American Beetles vol. 2, Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. pp. 537–539. ISBN 0-8493-0954-9.
White, Richard E. (1998) [1983]. A Field Guide to the Beetles of North America (Peterson Field Guides). Houghton Mifflin Harcourt. ISBN 0395910897.

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