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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Cladus: Macroheterocera
Superfamilia: Geometroidea

Familia: Geometridae
Subfamilia: Sterrhinae
Tribus: Scopulini
Genera: Aquilargilla – Dithalama – Isoplenodia – Lipomelia – Problepsis – Scopula – Somatina – Zythos
Source(s) of checklist:

Check (1): Lycaugidia

Additional references

Cui, L. & Jiang, N. 2018. Aquilargilla gen. nov., a new genus of Sterrhinae from China with description of two new species (Lepidoptera, Geometridae). Zootaxa 4514(3): 431–437. DOI: 10.11646/zootaxa.4514.3.8 Paywall Reference page.
Sihvonen, P. 2005: Phylogeny and classification of the Scopulini moths (Lepidoptera: Geometridae, Sterrhinae). Zoological journal of the Linnean Society 143(4): 473-530. DOI: 10.1111/j.1096-3642.2005.00153.x Reference page.
Sihvonen, P.; Kaila, L. 2004: Phylogeny and tribal classification of Sterrhinae with emphasis on delimiting Scopulini (Lepidoptera: Geometridae). Systematic entomology 29(3): 324-358. DOI: 10.1111/j.0307-6970.2004.00248.x Reference page.
Sihvonen, P.; Siljander, M. 2005: Species diversity and geographical distribution of Scopulini moths (Lepidoptera: Geometridae, Sterrhinae) on a world-wide scale. Biodiversity & conservation 14(3): 703-721. DOI: 10.1007/s10531-004-3921-8 Reference page.

Scopulini is a tribe of the geometer moth family (Geometridae), with about 900 species in seven genera. The tribe was described by Philogène Auguste Joseph Duponchel in 1845.


Scopulini as a family name is an old synonym of the subfamily Sterrhinae (Meyrick, 1892). The tribe Scopulini is divided into seven genera, of which only Scopula and Problepsis have species in Europe.

Scopulini Duponchel, 1845
Dithalama Meyrick, 1888 (4 species in Australia and Tasmania)
Isoplenodia Prout, 1932 (4 species in Africa)
Lipomelia Warren, 1893 (1 species from India to Taiwan)
Somatina Guenée, 1858 (44 species in Africa, East Asia and Australia)
Zythos D. S. Fletcher, 1979 (11 species from Indonesia up to Papua-New Guinea)
Problepsis Lederer, 1853 (51 species in the Palearctic, Africa, South-East Asia to Australia)
Scopula Schrank, 1802 (including Glossotrophia Prout, 1913 and Holarctias Prout, 1913) (over 800 species)


The phylogenetics of Scopulini was described in detail in 2005 by Pasi Sihvonen.


 Outgroup (Idaea)


 incertae sedis







 incertae sedis


Hausmann, Axel The Geometrid Moths of Europe, 2. Sterrhinae. Apollo Books, Stenstrup 2004, ISBN 87-88757-37-4
Abraham, D.; Ryrholm, N.; Wittzell, H.; Holloway, J. D.; Scoble, M. J.; Lofstedt, C.: "Molecular phylogeny of the subfamilies in Geometridae (Geometroidea: Lepidoptera)". Molecular Phylogenetics and Evolution. 20(1): 65-77 (2001)
Sihvonen, Pasi (April 1, 2005). "Phylogeny and classification of the Scopulini moths (Lepidoptera: Geometridae, Sterrhinae)". Zoological Journal of the Linnean Society. 143 (4): 473–530. doi:10.1111/j.1096-3642.2005.00153.x.

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