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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Lucanidae
Subfamilia: Aesalinae
Tribus: Aesalini - Ceratognathini - Nicagini
Overview of genera (9 + 3†)

Aesalus - Ceratognathus - Echinoaesalus - Hilophyllus - Holloceratognathus - Lucanobium - Mitophyllus - NicagusTrogellus – †Cretaesalus – †Juraesalus – †Sinaesalus


Aesalinae MacLeay, 1819

Type genus: Aesalus Fabricius, 1801




Holloway, B.A. 1998: A re-evaluation of the genera of New Zealand aesaline stag beetles (Coleoptera: Lucanidae). Journal of the Royal Society of New Zealand, 28(4): 641–656. DOI: 10.1080/03014223.1998.9517578 PDF
Holloway, B.A. 2007: Lucanidae (Insecta: Coleoptera). Fauna of New Zealand, (61)
Howden, H.F. & Lawrence, J.F. 1974: The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera, Lucanidae). Canadian journal of zoology 52: 1505-1510. DOI: 10.1139/z74-193 Open access. Reference page
Nikolajev, G.V.; Wang, B.; Liu, Y.; Zhang, H-c. 2011: Stag beetles from the Mesozoic of Inner Mongolia, China (Scarabaeoidea: Lucanidae). Acta palaeontologica sinica, 50(1): 41–47. [Publication date: 'March, 2011', sourced from first page of article]
Smith, A.B.T. 2006: A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society monograph, 5: 144–204.PDF


Tree of Life Web Project. 2007. Aesalinae. Version 02 May 2007 (temporary) in The Tree of Life Web Project
Tree of Life Web Project. 2007. Nicaginae. Version 02 May 2007 (temporary) in The Tree of Life Web Project

Vernacular names
日本語: マダラクワガタ亜科

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