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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Geotrupidae
Subfamilia: Bolboceratinae
Tribus: Athyreini
Genus: Athyreus
Species (47): A. acuticornis – A. aeneus – A. alvarengai – A. anneae – A. armatus – A. bellator – A. biarmatus – A. bicornus – A. bifurcatus – A. bilbergi – A. bilobus – A. brasilicus – A. capricornis – A. chalybeatus – A. championi – A. conspicuus – A. cyanescens – A. forcipatus – A. fritzi – A. gigas – A. giuglarisi – A. gulesseriani – A. hastifer – A. hemisphaericus – A. horatioi – A. howdeni – A. hypocritus – A. juanae – A. langeri – A. larseni – A. martinezi – A. mouraensis – A. nebulosus – A. nitidus – A. parvus – A. peckorum – A. pyriformis – A. soesilae – A. subarmatus – A. thilliezi – A. tribuliformis – A. tridens – A. tuberifer – A. unicornis – A. vavini – A. vicinus – A. zischkai

Athyreus MacLeay, 1819
Additional references

Boilly, O. 2012: Descriptions de deux nouveaux Athyreus MacLeay, 1819 (Coleoptera, Scarabaeoidea, Geotrupidae). Bulletin de la Société entomologique de France, 117(2): 193–197.
Howden, H.F. 1999: New Species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae). Coleopterists bulletin, 53: 339–354. JSTOR
Kohlmann, B.; Solís, A. 2012: New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama. Zootaxa, 3193: 28–52. Preview PDF
Makhan, D. 2008: Athyreus soesilae sp. nov., a new geotrupid beetle from Suriname (Coleoptera: Scarabaeoidea: Geotrupidae: Athyreini). Calodema supplementary paper, (82): 1–3.


Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on April 21, 2017]

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