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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Biblidinae
Tribus: Biblidini - Callicorini - Epicaliini - Epiphilini - Eubagini


Biblidinae Boisduval, 1833

Costa M., Viloria, A.L., Attal, S. & Orellana, A.M. 2014. Lepidoptera del Pantepui. Parte II. Description de nuevos Nymphalidae (Papilionoidea). Bulletin de la Société entomologique de France 119(1): 39–52. Reference page.
Wahlberg, N., Weingartner, E. & Nylin, S. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular phylogenetics and evolution 28(3): 473–484. DOI: 10.1016/S1055-7903(03)00052-6 PDF. Reference page.

Vernacular names
日本語: カバタテハ亜科
中文: 苾蛺蝶亞科

Biblidinae is a subfamily of nymphalid butterflies that includes the tropical brushfoots. This subfamily was sometimes merged within the Limenitidinae, but they are now recognized as quite distinct lineages. In older literature, this subfamily is sometimes called Eurytelinae.

As of 2008, some 340 valid species are in this subfamily, placed in 38[verification needed] genera. Most species of Biblidinae are Neotropical, but there are some Old World species and genera in the tribes Biblidini and Epicaliini.


The Biblidinae are a taxonomically stable monophyletic group, at least since the "wastebin genus" Catagramma was dismantled around 1950.

The tribes, in the presumed phylogenetic sequence and with notable genera also listed here, are:[1]

Biblidini Boisduval, 1833

Biblis Fabricius, 1807 (= Zonaga)
Ariadne Horsfield, 1829 (= Ergolis)
Laringa Moore, 1901
Eurytela Boisduval, 1833
Neptidopsis Aurivillius, 1898
Mesoxantha Aurivillius, 1898
Byblia Hübner, 1819 (= Hypanis Boisduval, 1833 (non Pander in Menetries, 1832: preoccupied))
Mestra Hübner, 1825 (= Cystineura)
Archimestra Munroe, 1949
Vila Kirby, 1871 (= Lonia, Neptis Hübner, 1819 (non Fabricius, 1807: preoccupied), Olina Doubleday, 1848 (non Robineau-Desvoidy, 1830: preoccupied))

Epicaliini Guenée, 1865

Catonephele Hübner, [1819]
Nessaea Hübner, [1819]
Myscelia Doubleday, 1844 (= Sagaritis Hübner, 1821 (non Billberg, 1820: preoccupied), Sea)
Sevenia Koçak, 1996 (= Crenis Boisduval, 1833 (non Hübner, 1821: preoccupied), Sallya Hemming, 1964 (non Yochelson, 1956: preoccupied))
Eunica Hübner, [1819]
Cybdelis Boisduval, 1836

Ageroniini Doubleday, 1847

Hamadryas – cracker butterflies, calico butterflies
Ectima Doubleday, 1848
Panacea Godman & Salvin, 1883 (= Pandora Doubleday, 1848 (non Lamarck, 1799: preoccupied))
Batesia Felder & Felder, 1862

Epiphilini Jenkins, 1987

Asterope Hübner, [1819]
Pyrrhogyra Hübner, 1819 (= Corybas)
Epiphile Doubleday, 1844
Lucinia Hübner, 1823 (= Autodea)
Bolboneura Godman & Salvin, 1877
Temenis Hübner, 1819 (= Callicorina, Paromia Hewitson, 1861 (non Westwood, 1851: preoccupied))
Nica Hübner, 1826 (= Pseudonica)
Peria Kirby, 1871 (= Pelia Doubleday, 1849 (non Bell, 1836: preoccupied))

Eubagini Burmeister, 1878

Dynamine Hübner, 1819 (= Arisba, Eubagis, Sironia)

Callicorini Orfila, 1952 – eighty-eights and relatives

Diaethria Billberg, 1820
Callicore Hübner, [1819]
Perisama Doubleday, 1849 (= Orophila)
Antigonis Felder, 1861 (= Lincoya)
Cyclogramma Doubleday 1847
Haematera Doubleday, 1849 (= Callidula)
Paulogramma Dillon, 1948
Catacore Dillon, 1948


The higher classification of Nymphalidae, at


Hill, R. I., Penz, C. M., & DeVries, P. J. (2002). Phylogenetic analysis and review of Panacea and Batesia butterflies (Nymphalidae). Journal of the Lepidopterists' Society, 56, 199-215.
Jenkins, D. W. (1983). Neotropical Nymphalidae. I. Revision of Hamadryas. Bulletin of the Allyn Museum, 81, 1-146.
Jenkins, D. W. (1984). Neotropical Nymphalidae. II. Revision of Myscelia. Bulletin of the Allyn Museum (Gainesville), 87, 1-64.
Jenkins, D. W. (1985). Neotropical Nymphalidae. III. Revision of Catonephele. Bulletin of the Allyn Museum of Entomology, 92, 1-65.
Jenkins, D. W. (1985). Neotropical Nymphalidae. IV. Revision of Ectima. Bulletin of the Allyn Museum (Gainesville), 95, 1-30.
Jenkins, D. W. (1986). Neotropical Nymphalidae. V. Revision of Epiphile. Bulletin of the Allyn Museum (Gainesville), 101, 1-70.
Jenkins, D. W. (1987). Neotropical Nymphalidae. VI. Revision of Asterope (=Callithea Auct.). Bulletin of the Allyn Museum of Entomology, 114, 1-66.
Jenkins, D. W. (1989). Neotropical Nymphalidae. VII. Revision of Nessaea. Bulletin of the Allyn Museum (Gainesville), 125, 1-38.
Jenkins, D. W. (1990). Neotropical Nymphalidae. VIII. Revision of Eunica. Bulletin of the Allyn Museum, 131, 1-177.

External links

The higher classification of Nymphalidae, at
Biblidinae Boisduval, 1833 at Markku Savela's Lepidoptera and Some Other Life Forms: Preliminary species list. Version of 18 March 2007. Retrieved 30 May 2007.
Pteron Misspelled as Biblinae. Images. In Japanese but with binomial names.

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