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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Dynastinae
Tribus: Oryctoderini
Genus: Chalcasthenes
Species (4): C. divinus – C. pulcher – C. squamigera – C. styracoceros

Chalcasthenes Arrow, 1937: 42

Type species: Chalcasthenes pulcher Arrow, 1937 (pp. 42–43) by monotypy.


Strehlia Frey, 1969: 480 [syn. by Jameson & Ratcliffe, 2009]
Type species: Strehlia squamigera Frey, 1969 (p. 481, f. 2) by monotypy.

Primary references

Arrow, G.J. 1937. Systematic notes on beetles of the subfamily Dynastinae with descriptions of a few species in the British Museum collection. Transactions of the Entomological Society of London 86: 35–58. [original description: p. 42]

Additional references

Jameson, M.L.; Ratcliffe, B.C. 2009: Revision of the genus Chalcasthenes Arrow (Coleoptera: Scarabaeidae: Dynastinae: Oryctoderini) from the Solomon Islands. Australian journal of entomology 48: 149–163.


Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on Jun 9, 2017]

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