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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Cyrestinae
Tribus: Cyrestini - Pseudergolini


Cyrestinae Guenée, 1865

Wahlberg, N., Weingartner, E. & Nylin, S. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular phylogenetics and evolution 28(3): 473–484. DOI: 10.1016/S1055-7903(03)00052-6 PDF. Reference page.

Vernacular names
日本語: イシガケチョウ亜科

Cyrestinae is the name of a small subfamily of nymphalid brush-footed butterflies. It is considered to include only three genera - Marpesia, Chersonesia, and Cyrestis - distributed in the tropics.[1]

The circumscription of the Cyrestinae has seen some changes in the recent years, when the former tribes Cyrestini and Pseudergolini were suggested to form a monophyletic clade, and the name was given to the proposed new subfamily,[2] but the tribes were split again later—as two independent subfamilies—as their positions within the Nymphalidae were defined more clearly.[1] It is now considered to be the sister group to the larger subfamily Nymphalinae, and only three genera are included:


Marpesia is Neotropical and Cyrestis and Chersonesia are mainly Oriental with a few species in the Afrotropics.[3]

N. Wahlberg , J. Leneveu , U. Kodandaramaiah , C. Peña , S. Nylin , A. V. L. Freitas , and A. V. Z. Brower (2009). Nymphalid butterflies diversify following near demise at the Cretaceous/Tertiary boundary Proceedings of the Royal Society of London Series B Biological Sciences 276: 4295–4302.
Niklas Wahlberg , Elisabet Weingartner , and Sören Nylin (2003). Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea) Molecular Phylogenetics and Evolution 28: 473–484.
Wahlberg, Niklas and Andrew V. Z. Brower. (2012). Cyrestinae Guenée 1865. Version 6 August 2012 (temporary). in The Tree of Life Web Project,

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