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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Elateriformia
Superfamilia: Byrrhoidea

Familia: Heteroceridae
Subfamilia: Heterocerinae
Tribus: Heterocerini
Genus: Heterocerus
Species: H. aragonicus – H. dayremi – H. debilipes – H. etruscus – H. ezeri – H. fausti – H. fenestratus – H. fijiensis – H. flexuosus – H. flindersi – H. fossor – H. fulvipes – H. fusculus – H. hankae – H. holosericeus – H. humilis – H. infuscatus – H. jelineki – H. jonasi – H. kamtschaticus – H. kaszabi – H. largsensis – H. lorenzevae – H. magnus – H. marginatus – H. mastersi – H. medius – H. morgani – H. mus – H. nepalensis – H. novaeselandiae – H. obsoletus – H. ottomerkli – H. pallidivestis – H. parallelus – H. persicus – H. prepsli – H. pulchellus – H. sauteri – H. scabriusculus – H. sichuanensis – H. simillimus – H. stultus – H. subpilosus – H. thebaicus – H. thieleni

Heterocerus Fabricius, 1792

Charpentier, R. 1968: A monograph of the family Heteroceridae (Coleoptera) of the Notogean region. Arkiv fuer zoologi, 20: 205–241.
Mascagni, A. 2020. Two new species of Heterocerus FABRICIUS, 1792 from Mexico (Coleoptera: Heteroceridae). Koleopterologische Rundschau 90: 253–256. Full article (PDF). Reference page.
Sazhnev, A.S. 2019. A new synonymy of Heterocerus fenestratus (Thunberg, 1784) (Coleoptera: Heteroceridae) and its first records for South Hemisphere. Zootaxa 4624(4): 589–592. DOI: 10.11646/zootaxa.4624.4.10. Reference page.
Sazhnev, A.S. 2018. Redescription of Heterocerus fausti Reitter, 1879, bona species (Coleoptera, Heteroceridae). Zootaxa 4441(3): 597–600. DOI: 10.11646/zootaxa.4441.3.12 Paywall Reference page.


Atlas of the family Heteroceridae of Russia

Heterocerus is a genus of beetles in the family Heteroceridae. There are at least 20 described species in Heterocerus.[1][2]
Heterocerus mollinus


Heterocerus angustatus Chevrolat, 1864
Heterocerus aptus Miller, 1994
Heterocerus brunneus Melsheimer, 1844
Heterocerus cinctus Motschulsky, 1858
Heterocerus coheni Skalický, 2007
Heterocerus collaris Kiesenwetter, 1851
Heterocerus crossi Miller, 1995
Heterocerus fenestratus (Thunberg, 1784)
Heterocerus inciertus (Pacheco, 1964)
Heterocerus infrequens Miller, 1994
Heterocerus insolens Miller, 1994
Heterocerus intermuralis Pacheco, 1963
Heterocerus mexicanus Sharp, 1882
Heterocerus mirus Miller, 1994
Heterocerus mollinus Kiesenwetter, 1851
Heterocerus pallidus Say, 1823
Heterocerus parrotus (Pacheco, 1964)
Heterocerus subtilis Miller, 1988
Heterocerus tenuis Miller, 1988
Heterocerus texanus (Pacheco, 1964)
Heterocerus undatus Melsheimer, 1844
Heterocerus unicus Miller, 1988
Heterocerus unituberculosus Miller, 1995
Heterocerus virginiensis Skalický, 2007


"Heterocerus Genus Information". Iowa State University. Retrieved 26 January 2018.

"Heterocerus Report". Integrated Taxonomic Information System. Retrieved 26 January 2018.

Pacheco, Francisco (1978). "Family: Heteroceridae". A Catalog of the Coleoptera of America North of Mexico, x + 8.

Further reading

Arnett, R. H. Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). (21 June 2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press LLC, Boca Raton, Florida ISBN 978-0-8493-0954-0.
Arnett, Ross H. (2000). American Insects: A Handbook of the Insects of America North of Mexico. CRC Press.
Richard E. White. (1983). Peterson Field Guides: Beetles. Houghton Mifflin Company.

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