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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Nymphalinae
Tribus: Kallimini
Genus: Kallima
Species: K. albofasciata – K. alompra – K. buxtoni – K. cymodoce – K. horsfieldi – K. inachus – K. knyvettii – K. limborgii – K. paralekta – K. philarchus – K. rumia – K. spiridiva

Kallima Doubleday, 1849

Orange Oakleaf (Kallima inachus) at Samsing, Duars, West Bengal W IMG 6241

Kallima inachus


Nakamura, N. & Wakahara, H., 2013: Notes on the butterflies of Laos (VIII): review of the genus Kallima Doubleday, [1849](Lepidoptera, Nymphalidae) with descriptions of two new subspecies. Butterflies (Teinopalpus), 63: 14–28.

Vernacular names
English: Leaf Butterflies

Kallima, known as the oakleaf or oak leaf butterflies, is a genus of butterflies of the subfamily Nymphalinae in the family Nymphalidae. They are found in east, south and southeast Asia. Their common name is a reference to the lower surface of their wings, which is various shades of brown like a dead leaf.

When the wings are held closed, this results in a remarkable masquerade of a dead leaf, further emphasized by their wing shape.[1]

This genus has traditionally also included a number of African species, but they are now usually placed in Kallimoides, Junonia (alternatively in Kamilla) and Mallika. The following species are currently members of the genus Kallima:[2][3]

Kallima albofasciata Moore, 1877 – Andaman oakleaf
Kallima alompra Moore, 1879 – scarce blue oak leaf
Kallima buxtoni Moore, 1879
Kallima horsfieldii (Kollar, 1844) – Sahyadri blue oakleaf[4]
Kallima inachus (Boisduval, 1846) – orange oakleaf, Indian oakleaf, dead leaf
Kallima limborgii Moore, 1879 – Peninsular Malaya leaf butterfly
Kallima knyvetti de Nicéville, 1886 – scarce blue oakleaf
Kallima paralekta (Horsfield, 1829) – Indian leafwing
Kallima philarchus (Westwood, 1848) – Ceylon blue oakleaf
Kallima spiridiva Grose-Smith, 1885


Cott, Hugh (1940). Adaptive Coloration in Animals. Oxford University Press. pp. 318–320.
"Kallima Doubleday, [1849]" at Markku Savela's Lepidoptera and Some Other Life Forms
Takashi Shirôzu & Akinori Nakanishi (1984). "A revision of the genus Kallima Doubleday (Lepidoptera, Nymphalidae)". Tyô to Ga. 34 (3): 97–110.
Valappil, B. and K. Saji. 2015. Kallima horsfieldii Kollar, 1844 – Blue Oakleaf. Kunte, K., P. Roy, S. Kalesh and U. Kodandaramaiah (eds.). Butterflies of India, v. 2.20. Indian Foundation for Butterflies.

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