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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Libytheinae
Genus: Libythea
Species: L. albonervulata – L. alompra – L. amaniana – L. antipoda – L. bardas – L. batchiana – L. celtis – L. cinyras – L. collenettei – L. geoffroy – L. labdaca – L. laius – L. lepita – L. myrrha – L. narina – L. orientalis

Libythea Fabricius, 1807

Type species: Papilio celtis (Laicharting, [1782])

Hecaerge Ochsenheimer, 1816
Chilea Billberg, 1820
Hypatus Hübner, 1822
Libythaeus Boitard, 1828
Dichora Scudder, 1889
Barbarothea Scudder, 1892


Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 284
Kawahara, A.Y. 2013. Systematic revision and review of the extant and fossil snout butterflies (Lepidoptera: Nymphalidae: Libytheinae). Zootaxa 3631(1): 001–074. DOI: 10.11646/zootaxa.3631.1.1 Preview. Reference page.
Samson, C., 1980: The butterflies of Santa Ana Island, with descriptions of new taxa from San Cristobal Island, Solomon Islands. Tyô to Ga 30 (3-4): 211–236. Abstract and full article: [1].

Vernacular names
English: Beaks; Snouts
中文: 喙蝶屬

Libythea is a widespread genus of nymphalid butterflies commonly called beaks or snouts. They are strong fliers and may even be migratory.[1]

Source: The higher classification of Nymphalidae, at
Note: Names preceded by an equal sign (=) are synonyms, homonyms, rejected names or invalid names.

Subfamily Libytheinae Boisduval, 1833

Libythea Fabricius, 1807 (= Hecaerge Ochsenheimer, 1816; = Chilea Billberg, 1820; = Hypatus Hübner, 1822; = Libythaeus Boitard, 1828; = Dichora Scudder, 1889)
Libythea geoffroy Godart, 1824
Libythea geoffroy geoffroy Godart, 1824
Libythea geoffroy alompra Moore, 1901 (= Libythea hauxwelli Moore, 1901)
Libythea geoffroy antipoda Boisduval, 1859 (= Libythea quadrinotata Butler, 1877)
Libythea geoffroy bardas Fruhstorfer, 1914
Libythea geoffroy batchiana Wallace, 1869
Libythea geoffroy celebensis Staudinger, 1859
Libythea geoffroy ceramensis Wallace, 1869
Libythea geoffroy deminuta Fruhstorfer, 1910
Libythea geoffroy genia Waterhouse, 1938
Libythea geoffroy howarthi Peterson, 1968
Libythea geoffroy maenia Fruhstorfer, 1902 (= Libythea geoffroy eugenia Fruhstorfer, 1910)
Libythea geoffroy nicevillei Olliff, 1891
Libythea geoffroy orientalis Godman & Salvin, 1888
Libythea geoffroy philippina Staudinger, 1889
Libythea geoffroy pulchra Butler, 1882 (= Libythea neopommerana Pagenstecher, 1896)
Libythea geoffroy sumbensis Pagenstecher, 1896
Libythea collenettei Poulton & Riley, 1928
Libythea narina Godart, 1819
Libythea narina narina Godart, 1819 (= Libythea hatami Kenrick, 1911)
Libythea narina canuleia Fruhstorfer, 1910
Libythea narina luzonica Semper, 1889
Libythea narina nahathaka Fruhstorfer, 1914
Libythea narina neratia Felder, 1864
Libythea narina rohini Marshall, 1880 (= Libythea libera de Niceville, 1890; = Libythea hybrida Martin, 1896; = Libythea tibera Pagenstecher, 1902)
Libythea narina sangha Fruhstorfer, 1914
Libythea narina sumbawana Fruhstorfer, 1914
Libythea labdaca Westwood & Hewitson, 1851
Libythea labdaca labdaca Westwood & Hewitson, 1851 (= Libythea labdaca werneri Fruhstrofer, 1903)
Libythea labdaca laius Trimen, 1879 (= Libythea labdaca cinyras Trimen, 1866; = Libythea labdaca lepitoides Moore, 1901)
Libythea ancoata Grose-Smith, 1891
Libythea tsiandava Grose-Smith, 1891
Libythea myrrha Godart, 1819
Libythea myrrha myrrha Godart, 1819
Libythea myrrha borneensis Fruhstorfer, 1914
Libythea myrrha carma Fruhstorfer, 1914
Libythea myrrha hecura Fruhstorfer, 1914
Libythea myrrha myrrhina Fruhstorfer, 1910
Libythea myrrha rama Moore, 1872
Libythea myrrha sanguinalis Fruhstorfer, 1898
Libythea myrrha thira Fruhstorfer, 1914
Libythea myrrha yawa Fruhstorfer, 1914
Libythea celtis (Fuessly, 1782) (original name = Papilio celtis Fuessly, 1782)
Libythea celtis celtis (Fuessly, 1782)
Libythea celtis amamiana Shirozu, 1956
Libythea celtis celtoides Fruhstorfer, 1909
Libythea celtis chinensis Fruhstorfer, 1909
Libythea celtis formosana Fruhstorfer, 1909
Libythea celtis lepita Moore, 1857
Libythea celtis sophene Fruhstorfer, 1914
Libythea cinyras Trimen, 1866
Libythea laius Trimen, 1879
Libythea laius laius Trimen, 1879
Libythea laius lepitoides Moore, 1903
Libythea laius tsiandava Grose-Smith 1891


Kawahara, A.Y. (2013). "Systematic revision and review of the extant and fossil snout butterflies (Lepidoptera: Nymphalidae: Libytheinae)". Zootaxa. 3631 (1): 1–74. doi:10.11646/zootaxa.3631.1.1.

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