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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Passalidae
Subfamilia: Incertae Sedis
Genus: Ophrygonius

Kon, M.; Johki, Y. 1991: A new species of Ophrygonius (Passalidae, Coleoptera) from Mt. Kinabalu, Sabah, Borneo. Japanese journal of entomology, 59 (3): 505–508. PDF
Kon, M., Johki, Y. & Araya, K. 2002. Notes on Ophrygonius emas (Iwase, 1998), comb. nov. (Coleoptera, Passalidae). Elytra 30(2): 451–453. PDF. Reference page.


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