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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Formicoidea

Subfamilia: Leptanillinae
Tribus: Anomalomyrmini
Genus: Protanilla
Species (9): P. bicolor – P. concolor – P. furcomandibula – P. gengma – P. jongi – P. lini – P. rafflesi – P. schoedli – P. tibeta – P. wardi

Protanilla Taylor in Bolton, 1990

Type species: Protanilla rafflesi Taylor, in Bolton, 1990

Baroni Urbani, C.; de Andrade, M.L. 2006: A new Protanilla Taylor, 1990 (Hymenoptera: Formicidae: Leptanillinae) from Sri Lanka. Myrmecologische Nachrichten, 8: 45–47. PDF
Bharti, H.; Akbar, S.A. 2015: First record of genus Protanilla (Hymenoptera: Formicidae: Leptanillinae) from India with description of a new species. Journal of Asia-Pacific entomology, 18(3): 573–576. Full article (PDF). Reference page.
Bolton, B. 1990: The higher classification of the ant subfamily Leptanillinae (Hymenoptera: Formicidae). Systematic entomology, 15 (3): 267–282. DOI: 10.1111/j.1365-3113.1990.tb00063.x PDF
Borowiec, M.L.; Schulz, A.; Alpert, G.D.; Baňař, P. 2011: Discovery of the worker caste and descriptions of two new species of Anomalomyrma (Hymenoptera: Formicidae: Leptanillinae) with unique abdominal morphology. Zootaxa, 2810: 1–14. Preview
Hsu, P-W., Hsu, F-C., Hsaio, Y. & Lin, C-C. 2017. Taxonomic notes on the genus Protanilla (Hymenoptera: Formicidae: Leptanillinae) from Taiwan. Zootaxa 4268(1): 117–130. DOI: 10.11646/zootaxa.4268.1.7. Reference page.
Terayama, M. 2009: A synopsis of the family Formicidae of Taiwan (Insecta, Hymenoptera). Research bulletin of Kanto Gakuen University Liberal Arts, 17: 81–266. [not seen]
Xu, Z.-H. 2002: A systematic study on the ant subfamily Leptanillinae of China (Hymenoptera: Formicidae). Acta Entomologica sinica 45(1): 115–120. Full article (PDF). Reference page.
Xu, Z.-H. 2012a: Furcotanilla, a New Genus of the Ant Subfamily Leptanillinae from China with Descriptions of Two New Species of Protanilla and P. rafflesi Taylor (Hymenoptera: Formicidae). Sociobiology 59(2): 477-491. Full article (PDF) on AntWiki Reference page.
Xu, Z.-H. & Zhang, J.L. 2002: Two new species of the ant subfamily Leptanillinae from Yunnan, China (Hymenoptera: Formicidae). Acta zootaxonomica sinica 27(1): 139–144. Full article (PDF). Reference page.


EoL [needs work]
Nomenclator Zoologicus

Protanilla is a genus of subterranean ants in the subfamily Leptanillinae.[2] Known from the Indomalayan realm, the genus contains about thirteen species. The genus was erected by Taylor (1990) for the type species P. rafflesi, described from workers from Peninsular Malaysia. Species in this genus have long and downcurved mandibles with peg-like tooth on the inner margins. Four species are known from China (P. bicolor, P. concolor, P. gengma and P. tibeta), one from Taiwan (P. lini), one from Sri Lanka (P. schoedli)[3][4] and a couple from India (P. wardi, P. flamma).[5][6]

Protanilla beijingensis Man et al., 2017
Protanilla bicolor Xu, 2002
Protanilla concolor Xu, 2002
Protanilla flamma Baidya & Bagchi, 2020
Protanilla furcomandibula Xu & Zhang, 2002
Protanilla gengma Xu, 2012
Protanilla izanagi Terayama, 2013
Protanilla jongi Hsu et al., 2017
Protanilla lini Terayama, 2009
Protanilla rafflesi Taylor, 1990
Protanilla schoedli Baroni Urbani & De Andrade, 2006
Protanilla tibeta Xu, 2012
Protanilla wardi Bharti & Akbar, 2015


Bolton, B. (2021). "Protanilla". AntCat. Retrieved 2 December 2021.
"Genus: Protanilla". AntWeb. Retrieved 21 September 2013.
Borowiec, M. L.; Schulz, A.; Alpert, G. D.; Baňař, P. (2011). "Discovery of the worker caste and descriptions of two new species of Anomalomyrma (Hymenoptera: Formicidae: Leptanillinae) with unique abdominal morphology". Zootaxa. 2810: 1–14. doi:10.11646/zootaxa.2810.1.1.
Xu, Z.-H. (2012). "Furcotanilla, a new genus of the ant subfamily Leptanillinae from China with descriptions of two new species of Protanilla and P. rafflesi Taylor (Hymenoptera: Formicidae)". Sociobiology. 59 (2): 477–491. doi:10.13102/sociobiology.v59i2.612.
Bharti, H.; Akbar, S.A. (2015). "First record of genus Protanilla (Hymenoptera: Formicidae: Leptanillinae) from India with description of a new species". Journal of Asia-Pacific Entomology. Elsevier, Korean Society of Applied Entomology and the Taiwan Entomological Society. 18 (3): 573–576. doi:10.1016/j.aspen.2015.04.008.
Baidya, Pronoy; Bagchi, Sumanta (2020). "A new species of Protanilla Taylor 1990 (Hymenoptera: Formicidae: Leptanillinae) from India". Halteres. 11: 19–24. doi:10.5281/zenodo.3941686.

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