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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Aphodiinae
Tribus: Psammodiini
Subtribus (2): Psammodiina - Rhyssemina

Psammodiini Mulsant, 1842

Pittino, R. 2006: Redescription of the genus Petrovitzius Rakovič, 1979, and descriptions of Rakovicius gen. nov. from East Asia, and Rakovicius kawaii sp. nov. from Taiwan (Coleoptera, Aphodiidae, Psammodiinae). Kogane, (7 ): 11–21. [Publication date: 'Jul. 2006', sourced from first page of article]
Smith, A.B.T. 2006: A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society monograph, 5: 144–204. DOI: 10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2 PDF
Stebnicka, Z.T.; Howden, H.F. 1996: Australian genera and species in the tribes Odontolochini, Psammodiini, Rhyparini, Stereomerini and part of the Eupariini (Coleoptera: Scarabaeoidea: Aphodiinae). Invertebrate taxonomy, 10 (1): 97–170. DOI: 10.1071/IT9960097

Psammodiini is a tribe of aphodiine dung beetles in the family Scarabaeidae. There are about 12 genera and at least 50 described species in Psammodiini.[1][2]
Tesarius caelatus

Diastictus Mulsant, 1842
Geopsammodius Gordon & Pittino, 1992
Leiopsammodius Rakovic, 1981
Neopsammodius Rakovic, 1986
Odontopsammodius Gordon & Pittino, 1992
Parapsammodius Gordon & Pittino, 1992
Platytomus Mulsant, 1842
Pleurophorus Mulsant, 1842
Psammodius Fallén, 1807
Rhyssemus Mulsant, 1842
Tesarius Rakovic, 1981
Trichiorhyssemus Clouët, 1901


"Psammodiini Tribe Information". Retrieved 1 February 2018.

"Psammodiini Report". Integrated Taxonomic Information System. Retrieved 1 February 2018.

Bouchard, P., Y. Bousquet, A. Davies, M. Alonso-Zarazaga, J. Lawrence, C. Lyal, A. Newton, et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys, vol. 88, 1–972.

Further reading

Arnett, R. H. Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). (21 June 2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press LLC, Boca Raton, Florida ISBN 978-0-8493-0954-0.
Arnett, Ross H. (2000). American Insects: A Handbook of the Insects of America North of Mexico. CRC Press.
Richard E. White. (1983). Peterson Field Guides: Beetles. Houghton Mifflin Company.

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