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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Scarabaeini
Genera (8): DrepanopodusEscarabaeusKheperPachylomeraPachysomaScarabaeolusScarabaeusSceliages


Scarabaeini Latreille, 1802

Pachysomatini Ferreira, 1953
Actinophorini Ádám, 2003

Type genus

Actinophorini Ádám: Actinophorus Creutzer, 1799 [=Scarabaeus Linnaeus, 1758] [stem = Actinophor-]
Pachysomatini Ferreira: Pachysoma MacLeay, 1821 [stem = Pachysomat-]
Scarabaeini Latreille: Scarabaeus Linnaeus, 1758 [stem = Scarabae-]

cited sources

Forgie, S.A.; Kryger, U.; Bloomer, P.; Scholtz, C.H. 2006: Evolutionary relationships among the Scarabaeini (Coleoptera: Scarabaeidae) based on combined molecular and morphological data. Molecular phylogenetics and evolution, 40 (3): 662–678. DOI: 10.1016/j.ympev.2006.04.018
Latreille, 1802:
Linnaeus, C. 1758: Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Laurentii Salvii, Stockholm. BHL

Additional references

Forgie, S.A.; Philips, T.K.; Scholtz, C.H. 2005: Evolution of the Scarabaeini (Scarabaeidae: Scarabaeinae). Systematic entomology 30(1): 60–96. DOI: 10.1111/j.1365-3113.2004.00273.x Reference page.
Mostert, L.E. & Holm, E. 1982. Notes on the flightless Scarabaeina (Coleoptera: Scarabaeidae) with a description of a new species. Cimbebasia (A) 5: 273–284. Reference page.
Mostert, L.E.; Scholtz, C.H. 1986: Systematics of the subtribe Scarabaeina (Coleoptera: Scarabaeidae). Entomology memoir, Department of Agriculture and Water Supply, Republic of South Africa, (65 ) [not seen]
Smith, A.B.T. 2006: A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society monograph, 5: 144–204. DOI: 10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2 PDF
Sole, C.L.; Wirta, H.; Forgie, S.A.; Scholtz, C.H. 2011: Origin of Madagascan Scarabaeini dung beetles (Coleoptera: Scarabaeidae): dispersal from Africa. Insect systematics & evolution, 42 (1): 29–40. DOI: 10.1163/187631211X552800

unpublished theses/reports

Forgie, S.A. 2003: Phylogeny of the Scarabaeini (Coleoptera: Scarabaeidae). PhD Thesis. University of Pretoria, South Africa.


Tribe Scarabaeini (Scarabaeidae: Scarabaeinae) - Atlas of scarab beetles of Russia

The Scarabaeini[1] are a tribe of old-world dung beetle genera,[2] erected by Pierre André Latreille.
Pachylomera femoralis

BioLib[2] lists:

Allogymnopleurus Janssens, 1940
Garreta Janssens, 1940
Gymnopleurus Illiger, 1803
Mnematium MacLeay, 1821
Neosisyphus G. Müller, 1942
Nesosisyphus Vinson, 1946
Pachylomera Kirby, 1828
Paragymnopleurus Shipp, 1897
Scarabaeus Linnaeus, 1758
Sceliages Westwood, 1837
Sisyphus Latreille, 1807

Note: a number of taxa including Kheper and Mnematidium are now considered subgenera of Scarabaeus.

Latreille PA (1802) Histoire naturelle, générale et particulière des Crustacés et des Insectes. Tome 3. F. Dufart, Paris. tribus Scarabaeini Latreille, 1802 (retrieved 23 August 2021)

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