Gymnusa

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Staphylinoidea

Familia: Staphylinidae
Subfamilia: Aleocharinae
Tribus: Gymnusini
Genus: Gymnusa
Species (14): G. absens – G. anatolica – G. antiqua – G. atra – G. brevicollis – G. campbelli – G. grandiceps – G. inexspectata – G. konopackii – G. lindrothi – G. miyashitai – G. pseudovariegata – G. smetanai – G. variegata
Name

Gymnusa Gravenhorst, 1806: 173

Type species: Gymnusa sinuata Gravenhorst, 1806, fixed by monotypy.

Synonymy

Gymnusa Mannerheim, 1830: 66 [also 1831: 480; not as new, attributed to Karsten; synonymic homonym]
Ischnocephalus Gistel, 1856: 387 [without description]
Type species: Staphylinus brevicollis Paykull, 1800, fixed by monotypy.

References
Primary references

Gravenhorst, J.L.C. 1806. Monographia Coleopterorum Micropterorum. Gottingae: Henricus Dieterich, 236 + [12] pp. [original description: p. 173]

Additional references

Hammond, P. 1975: The phylogeny of a remarkable new genus and species of gymnusine staphylinid (Coleoptera) from the Auckland Islands. Journal of entomology (B), 44: 153–173. DOI: 10.1111/j.1365-3113.1975.tb00010.x
Klimaszewski, J. 1979: A revision of the Gymnusini and Deinopsini of the world. Coleoptera: Staphylinidae, Aleocharinae. Canada Department of Agriculture Research Branch monograph, (25).
Naomi, S.-I. 1994: Description of a new species of genus Gymnusa Gravenhorst (Coleoptera: Staphylinidae) from Hokkaido, Japan. Natural history research, 3(1): 47–50.

Gymnusa is a genus of rove beetles in the subfamily Aleocharinae and tribe Gymnusini. The coloration of this genus is highly coherent, with most species black and a few dark, dark brown. They range in length from 4.2 millimeters to 6.5 millimeters.[1]

Gymnusa live in semi-aquatic and aquatic environments, but never marine: they live in detritus in swamps, in bogs, streams, and slow-moving rivers. They live in the Nearctic and Palearctic in Europe and North America.[1]
References

"Gymnusa". 2001. Archived from the original on April 2, 2015. Retrieved February 28, 2015.