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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Staphylinoidea
Familiae (6): Agyrtidae - Hydraenidae - Leiodidae - Ptiliidae - Silphidae - Staphylinidae




Gerlach, J. 2009: Superfamily Staphylinoidea. Pp. 56-78 in: Gerlach, J. (ed.) The Coleoptera of the Seychelles Islands. Pensoft Publishers, Sofia-Moscow. ISBN 9789546424983
Grebennikov, V.V.; Newton, A.F. 2012: Detecting the basal dichotomies in the monophylum of carrion and rove beetles (Insecta: Coleoptera: Silphidae and Staphylinidae) with emphasis on the Oxyteline group of subfamilies. Arthropod systematics & phylogeny, 70(3): 133–165. full article (PDF)
Newton, A.F., jr. 1985: South temperate Staphylinoidea (Coleoptera): their potential for biogeographic analysis of austral disjunctions. Pp. 180-220 in Ball, G.E. (ed.) Taxonomy, phylogeny and zoogeography of beetles and ants. Series entomologica (Dordrecht), 33.
Atlas of beetles of Russia: the superfamily STAPHYLINOIDEA
Catalogue of Palearctic Coleoptera. Vol. 2 ed. I. Lobl, & A. Smetana, Apollo Books, Stenstrup, Denmark, 2004
ISBN 87-88757-74-9, p.102


Australian Faunal Directory
PEET - Austral Staphylinidae

Vernacular names
català: Estafilinoïdeu
suomi: Haiskiaismaiset
magyar: Holyvaszerűek
한국어: 반날개상과
norsk: Åtsel- og rovbiller
русский: Стафилиноидные
українська: Стафіліноїдні
Tiếng Việt: Liên họ Cánh cụt
中文: 隐翅虫总科

Staphylinoidea is a superfamily of beetles. It is a very large and diverse group with worldwide distribution.

Most staphylinoids are middle-sized or small beetles with a pair of shortened elytra. The hind wings have no accessory posterior ridge (locking device), no wedge cell and apical cell. Abdominal 8th segment not entirely invaginated in the 7th. Head mostly without coronal suture.[1]
Systematics and evolution

Staphylinoidea contains the following subgroups:[2][3][4][5]

Family Agyrtidae C.G. Thomson 1859 (primitive carrion beetles)
Subfamily Agyrtinae Thomson, 1859
Subfamily Necrophilinae Newton, 1997
Subfamily Pterolomatinae Thomson, 1862
Family Hydraenidae Mulsant 1844 (minute moss beetles)
Subfamily Hydraeninae Mulsant, 1844
Subfamily Ochthebiinae C. G. Thomson, 1860
Subfamily Orchymontiinae Perkins, 1997
Subfamily Prosthetopinae Perkins, 1994
Family Leiodidae Fleming 1821 (round fungus beetles)
Subfamily Camiarinae Jeannel, 1911
Subfamily Catopocerinae Hatch, 1927
Subfamily Cholevinae Kirby, 1837
Subfamily Coloninae Horn, 1880
Subfamily Leiodinae Fleming, 1821
Subfamily Platypsyllinae Ritsema, 1869
Family Ptiliidae Erichson 1845 (featherwing beetles)
Subfamily Acrotrichinae Reitter, 1909
Subfamily Cephaloplectinae Sharp, 1883
Subfamily Nanosellinae Barber, 1924
Subfamily Ptiliinae Erichson, 1845
Family Silphidae Latreille 1807 (carrion beetles)
Subfamily Nicrophorinae Kirby, 1837
Subfamily Silphinae Latreille, 1806
Family Staphylinidae Latreille 1802 (rove beetles)
Subfamily Aleocharinae Fleming, 1821
Subfamily Apateticinae Fauvel, 1895
Subfamily Dasycerinae Reitter, 1887
Subfamily Empelinae Newton and Thayer, 1992
Subfamily Euaesthetinae Thomson, 1859
Subfamily Glypholomatinae Jeannel, 1962
Subfamily Habrocerinae Mulsant and Rey, 1876
Subfamily Leptotyphlinae Fauvel, 1874
Subfamily Megalopsidiinae Leng, 1920
Subfamily Micropeplinae Leach, 1815
Subfamily Microsilphinae Crowson, 1950
Subfamily Neophoninae Fauvel, 1905
Subfamily Olisthaerinae Thomson, 1858
Subfamily Omaliinae MacLeay, 1825
Subfamily Osoriinae Erichson, 1839
Subfamily Oxyporinae Fleming, 1821
Subfamily Oxytelinae Fleming, 1821
Subfamily Paederinae Fleming, 1821
Subfamily Phloeocharinae Erichson, 1839
Subfamily Piestinae Erichson, 1839
Subfamily Proteininae Erichson, 1839
Subfamily Protopselaphinae Newton and Thayer, 1995
Subfamily Pselaphinae Latreille, 1802
Subfamily Pseudopsinae Ganglbauer, 1895
Subfamily Scaphidiinae Latreille, 1806
Subfamily Scydmaeninae Leach, 1815
Subfamily Solieriinae Newton and Thayer, 1992
Subfamily Staphylininae Latreille, 1802
Subfamily Steninae MacLeay, 1825
Subfamily Tachyporinae MacLeay, 1825
Subfamily Trichophyinae Thomson, 1858
Subfamily Trigonurinae Reiche, 1866
† Subfamily Protactinae Heer, 1847

The unambiguous fossil record dates back to Triassic, and an early Mesozoic origin of the group is probable.[6]

M.Hansen. Phylogeny and classification of the staphyliniform beetle families (Coleoptera). Biologiske Skrifter 48, Copenhagen, 1997
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
Beutel, R. G. and Leschen, R.A.B. 2005. Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults. Systematic Entomology
J. F. Lawrence and A. F. Newton, Jr. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). pp. 779-1006 In: J. Pakaluk & S. A. Slipinski (Eds.): Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa
"Superfamily Staphylinoidea - Rove, Carrion and Fungus Beetles". Retrieved 2019-06-16.
Grimaldi, D. and Engel, M.S. (2005). Evolution of the Insects. Cambridge University Press

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