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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Staphylinoidea

Familia: Hydraenidae
Subfamiliae (4): Hydraeninae - Ochthebiinae - Orchymontiinae - Prosthetopinae

Overview of genera

Davidraena – Gondraena – Menomadraena – Trinomadraena –

Hydraenidae Mulsant, 1844

Limnebiidae Mulsant, 1844


Template:Mulsant, 1844

Deler-Hernández, A.; Delgado, J.A. 2012: The Hydraenidae of Cuba (Insecta: Coleoptera) I: Contribution to the fauna of eastern Cuba. Zootaxa 3478: 213–238. Preview Reference page.
Deler-Hernández, A. & Delgado, J.A. 2017. The Hydraenidae of Cuba (Insecta: Coleoptera) II: Morphology of preimaginal stages of six species and notes on their biology. Zootaxa 4238(4): 451–498. DOI: 10.11646/zootaxa.4238.4.1. Reference page.
Ertorun, N. et al. 2011: Checklist of the Hydraenidae (Coleoptera) of Turkey, with notes on distribution. Zootaxa, 3055: 22–42. Preview
Hansen, M. 1998: Hydraenidae (Coleoptera). World catalogue of insects, 1. Stenstrup: Apollo Books. ISBN 8788757277
Jäch, M.A. 1994. Description of Gondraena gen.n. from South India and Madagascar and Davidraena gen.n. from South India (Hydraenidae). Koleopterologische Rundschau 64: 85-102. Full article (PDF). Reference page.
Jäch, M.A.; Madl, M. 2009: Water beetles of Seychelles (Coleoptera). Pp. 11-35, plates 1-2 in: Gerlach, J. (ed.) The Coleoptera of the Seychelles Islands. Pensoft Publishers, Sofia-Moscow. ISBN 9789546424983
Lee, D.-H. & Ahn, K.-J. 2016. A taxonomic review of the Hydraenidae in South Korea (Coleoptera). ZooKeys 634: 57–78. DOI: 10.3897/zookeys.634.10640 Open access. Reference page.
Perkins, P.D. 1997: Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of the Carnegie Museum, 66: 89–207.
Perkins, P.D. 2017. Hydraenidae of Madagascar (Insecta: Coleoptera). Zootaxa 4342(1): 1–264. DOI: 10.11646/zootaxa.4342.1.1. Reference page.
Salah, M.; Cueto, J.A.R.; Valladares, L.F. 2014: An annotated checklist of the aquatic Polyphaga (Coleoptera) of Egypt I. Family Hydraenidae. Zootaxa 3873(3): 275–285. DOI: 10.11646/zootaxa.3873.3.6 Reference page.
Catalogue of Palearctic Coleoptera. Vol. 2 ed. I. Lobl, & A. Smetana, Apollo Books, Stenstrup, Denmark, 2004
ISBN 87-88757-74-9, p.102

Vernacular names
čeština: Vodanovití
Deutsch: Langtasterwasserkäfer
English: Minute Moss Beetle
lietuvių: Prūdvabaliai
norsk: Palpebiller
русский: Водобродки

Hydraenidae is a family of very small aquatic beetles, sometimes called "minute moss beetles", with a worldwide distribution. These beetles are generally 1–3 mm in length (although some species reach 7 mm) with clubbed antennae. They do not swim well and are generally found crawling in marginal vegetation. Most are phytophagous but a few saprophagous and predatory species are known.

The family consists of more than 2000 species belonging to over 50 genera. Genus Hydraena alone includes nearly 900 described species worldwide.

The beetles are semi-aquatic, riparian and can walk on water surface. They feed on algae, bacteria, protozoans and detritus on wet stones and plant matter.

The larvae, although preferring damp habitats near water, are non-aquatic and tend to drown easily. They have a distinct labrum and well developed galea of maxillae. 9th abdominal tergum has a pair of apical, re-curved ventral hook and 2-segmented urogomphi.[1][2]

1 Genera
1.1 Extinct genera
2 References
3 External links

Ochthebius capicola

These 56 genera belong to the family Hydraenidae:[3][4][5]

Adelphydraena Perkins, 1989
Archaeodraena Jäch & Yamamoto, 2017
Aulacochthebius Kuwert, 1887
Calobius Wollaston, 1854
Cobalius Rey, 1886
Coelometopon Janssens, 1972
Davidraena Jäch, 1994
Decarthrocerus Orchymont, 1948
Discozantaena Perkins & Balfour-Browne, 1994
Edaphobates Jäch & Díaz, 2003-01
Enicocerus Stephens, 1829
Ginkgoscia Jäch & Díaz, 2004-01
Gondraena Jäch, 1994
Gymnanthelius Perkins, 1997
Gymnochthebius Orchymont, 1943
Haptaenida Perkins, 1997
Heptaenida Perkins, 1997
Homalaena Ordish, 1984
Hughleechia Perkins, 1981
Hydraena Kugelann, 1794
Hydraenida Germain, 1901
Hydroenida Germain, 1901
Hymenodes Mulsant, 1844
Laeliaena Sahlberg, 1900
Limnebius Leach, 1815
Madagaster Perkins, 1997
Menomadraena Perkins, 2017
Meropathus Enderlein, 1901
Micragasma Sahlberg, 1900
Neochthebius Orchymont, 1932
Nucleotops Perkins & Balfour-Browne, 1994
Ochtebiites Ponomarenko, 1977
Ochthebius Leach, 1815
Ochtheosus Perkins, 1997
Oomtelecopon Perkins, 2005
Orchymontia Broun, 1919
Parasthetops Perkins & Balfour-Browne, 1994-31
Parhydraena Orchymont, 1937
Parhydraenopsis Perkins, 2009-16
Phothydraena Kuwert, 1888
Pneuminion Perkins, 1997
Podaena Ordish, 1984
Prionochthebius Kuwert, 1887
Prosthetops Waterhouse, 1879
Protochthebius Perkins, 1997
Protosthetops Perkins, 1994
Protozantaena Perkins, 1997
Pterosthetops Perkins, 1994
Sebasthetops Jäch, 1998-01
Sicilicula Balfour-Browne, 1958
Spanglerina Perkins, 1980
Trinomadraena Perkins, 2017
Tympallopatrum Perkins, 1997
Mesoceration Janssens, 1967
Parhydraenida Balfour-Browne, 1975
Tympanogaster Janssens, 1967

Extinct genera

†Archaeodraena Jäch and Yamamoto 2017, Burmese amber, Myanmar, Cenomanian
†Ochtebiites Ponomarenko 1977 Abasheva Formation, Russia, Pliensbachian, Ichetuy Formation, Russia, Oxfordian, Karabastau Formation, Kazakhstan, Oxfordian, Mogotuin Formation, Mongolia, Aptian


Hydraenidae, Lucid Central.
L. Watson and M.J. Dallwitz. Hydraenidae, Insects of Britain and Ireland: the families of Coleoptera. Version: 7 October 2018.
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
"Hydraenidae". GBIF. Retrieved 16 June 2019.
"Hydraenidae Report". Integrated Taxonomic Information System. Retrieved 16 June 2019.

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