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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Heliconiinae
Tribus: Heliconiini
Genera: AgraulisDioneDryadulaDryasEueidesHeliconiusPhilaethriaPodotricha


Heliconiini Swainson, 1822

Brower, A.V.Z., 1994: Phylogeny of Heliconius butterflies inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 3: 159–174.
Brower, A.V.Z., 1994: Rapid morphological radiation and convergence among races of the butterfly Heliconius erato inferred from patterns of mitochondrial DNA evolution. Proceedings of the National Academy of Sciences USA 91: 6491–6495.
Brower, A.V.Z., 1996: Parallel race formation and the evolution of mimicry in Heliconius butterflies: a phylogenetic hypothesis from mitochondrial DNA sequences. Evolution 50: 195–221.
Brower, A.V.Z., 1997: The evolution of ecologically important characters in Heliconius butterflies (Lepidoptera: Nymphalidae): a cladistic review. Zoological Journal of the Linnean Society, 119: 457–472.
Brower, A.V.Z., and M.G. Egan, 1997: Cladistic analysis of Heliconius butterflies and relatives (Nymphalidae: Heliconiiti): a revised phylogenetic position for Eueides based on sequences from mtDNA and a nuclear gene. Proceedings of the Royal Society of London Series B Biological Sciences 264: 969–977.
Brown, K.S., 1973: The Heliconians of Brazil (Lepidoptera: Nymphalidae). Part V. Three new subspecies from the Mato Grosso and Rondonia. Bulletin of the Allyn Museum 13: 1–19. Full article: [1].
Brown Jr., K.S., 1981: The biology of Heliconius and related genera. Annual Review of Entomology 26: 427–456.
Emsley, M., 1963: A morphological study of imagine Heliconiinae (Lep.: Nymphalidae) with a consideration of the evolutionary relationships within the group. Zoologica 50: 191–254.
Kozak, K.M., Wahlberg, N., Neild, A.F.E., Dasmahapatra, K.K., Mallet, J., & Jiggins, C.D., 2015: Multilocus Species Trees Show the Recent Adaptive Radiation of the Mimetic Heliconius Butterflies. Systematic Biology 64(3): 505–524.
Lamas, G., (ed) 2004: Atlas of Neotropical Lepidoptera. Checklist: Part 4A, Hesperioidea - Papilionoidea. Scientific Publishers, Gainsville, Florida.
Michner, C. D., 1942: A generic revision of the Heliconiinae (Lepidoptera: Nymphalidae). American Museum Novitates 1197: 1–8.
Neukirchen, W.M., 2000: Two new subspecies of Ecuadorian Heliconiinae (Lepidoptera: Nymphalidae). Entomologische Zeitschrift 110 (5): 141–143.
Penz, C.M., 1999: Higher level phylogeny for the passion-vine butterflies (Nymphalidae, Heliconiinae) based on early stage and adult morphology. Zoological Journal of the Linnean Society, 127: 277–344.
Penz, C.M. & Peggie, D., 2003: Phylogenetic relationships among Heliconiinae genera based on morphology (Lepidoptera: Nymphalidae). Systematic Entomology 28: 451–479.
Ramos, R.R., Francini, R.B., Habib, M.E.E.M. & Freitas, A.V.L. 2021. Seasonal Patterns of Host Plant Use in an Assemblage of Heliconiini Butterflies (Lepidoptera: Nymphalidae) in a Neotropical forest. Neotropical Entomology 50(3): 358–365. DOI: 10.1007/s13744-021-00855-5.Paywall. Reference page. .

Heliconiini is a tribe of butterflies in the subfamily Heliconiinae, also known as the passion-vine butterflies. This group has roughly 100 species and subspecies distributed primarily in the Neotropics.[1][2][3]
Genera and select species

Agraulis (Boisduval & Le Conte, 1833) monotypic
Agraulis vanillae (Linnaeus, 1758) – Gulf fritillary
Dione (Hübner, 1819)
Dione juno (Cramer, 1779) – Juno silverspot, Juno longwing
Dryadula (Michener, 1942) monotypic
Dryadula phaetusa (Linnaeus, 1758) – banded orange, orange tiger
Dryas (Hübner, 1807) monotypic
Dryas iulia (Hübner, 1807) – Julia longwing
Eueides (Hübner, 1816) – longwings
Heliconius (Kluk, 1802) – brush-footed butterflies
Philaethria (Billberg, 1820)
Podotricha (Michener, 1942)

Two additional genera, Laparus Billberg, 1820 and Neruda Turner, 1976, have recently been synonymized with Heliconius.
See also

List of fritillaries (butterflies)


Tree of Life Web Project Archived 2010-07-11 at the Wayback Machine
Tribe Heliconiini at Markku Savela's Lepidoptera and Some Other Life Forms

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