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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Libytheinae
Genera: Libythea - Libytheana

Libytheinae Boisduval, 1833

Kawahara, Akito Y., 2006: Biology of the snout butterflies (Nymphalidae: Libytheinae) Part 1. Libythea Fabricius. Trans. lepid. Soc. Japan 57 (1): 13–33. [1]
Kawahara, A.Y. 2013. Systematic revision and review of the extant and fossil snout butterflies (Lepidoptera: Nymphalidae: Libytheinae). Zootaxa 3631(1): 001–074. DOI: 10.11646/zootaxa.3631.1.1 Preview. Reference page.

Vernacular names
日本語: テングチョウ亜科
中文: 喙蝶亞科

The Libytheinae are a nymphalid subfamily known as snout butterflies, containing two valid genera and about ten species: six in Libythea and four in Libytheana. The common name refers to the thick labial palps (pedipalps) that look like a "snout" in this subfamily. In older literature, this group was recognized as the family Libytheidae. They are medium-sized and typically a drab brown.[1] The front legs are reduced in length and the ventral hindwings are cryptically colored to help them blend in with their surroundings.[1] While at rest, the members of this subfamily keep their wings tightly closed to resemble dead leaves.[1]

Libytheinae is a subfamily of the family Nymphalidae:[2]

Family Nymphalidae Rafinesque, 1815
Subfamily Libytheinae Boisduval, 1833
Libythea Fabricius, 1807
Libytheana Michener, 1943


Daniels, Jaret C. (2003). Butterflies of the Carolinas. Cambridge, MN: Adventure Publications, Inc. p. 12. ISBN 1-59193-007-3.

The higher classification of Nymphalidae, at

Freitas, A. V. L., & Brown, K. S., Jr. (2004). Phylogeny of the Nymphalidae (Lepidoptera). Systematic Biology 53(3):363-383.
Kawahara, A. Y. (2003). Rediscovery of Libythea collenettei Poulton & Riley (Nymphalidae: Libytheinae) in the Marquesas, and a description of the male. Journal of the Lepidopterists' Society 57:81-85.
Shields, O. (1984). A revised, annotated checklist of world Libytheidae. Journal for Research on the Lepidoptera 22: 264-266.

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