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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Geotrupidae
Subfamilia: Bolboceratinae
Tribus: AthyreiniBolbelasminiBolboceratiniBolbochromini – Eubolbitini – Eucanthini – Gilletinini – Odonteini – Stenaspidiini

Genera (46 + 2†): †Amberathyreus – Athyreus – Australobolbus – BlackbolbusBlackburniumBolbafferBolbaffroidesBolbelasmus – Bolbobaineus – Bolbocaffer – Bolboceras – Bolbocerastes – Bolboceratex – Bolboceratops – Bolbocerodema – Bolboceroides – Bolbocerosoma – Bolbochromus – Bolbogonium – Bolbohamatum – Bolboleaus – Bolborhachium – Bolborhinum – Bolborhombus – Bolbothyreus – Bolbotrypes – Bradycinetulus – †Cretobolbus – Elephastomus – Eubolbitus – Eucanthus – Gilletinus – Halffterobolbus – Meridiobolbus – Mimobolbus – Namibiobolbus – Namibiotrupes – Neoathyreus – Odonteus – Parabolbapium – Parathyreus – Pereirabolbus – PrototrupesPseudoathyreusSocotrabolbusSomalobolbusStenaspidiusZefevazia


Bolboceratinae Mulsant, 1842

Bolboceratidae Mulsant, 1842 [status: Scholtz & Browne (1996: 613)]

Additional references

Bezděk, A. & Hájek, J. 2009. Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic - Scarabaeoidea: Bolboceratidae, Geotrupidae, Glaphyridae, Hybosoridae, Ochodaeidae and Trogidae. Acta Entomologica Musei Nationalis Pragae 49(1): 297–332. PDF Reference page.
Scholtz, C.H.; Browne, D.J. 1996: Polyphyly in the Geotrupidae (Coleoptera: Scarabaeoidea): a case for a new family. Journal of natural history, 30: 597–614. DOI: 10.1080/00222939600770311
Smith, A.B.T. 2006: A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Society monograph, 5: 144–204. DOI: 10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2 PDF


Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on April 22, 2017]

Bolboceratinae is a subfamily of earth-boring scarab beetles in the family Geotrupidae. There are about 8 genera and at least 40 described species in Bolboceratinae.[1][2][3]

Catalogue of Life[4] and GBIF[5] now consider Bolboceratinae to be a family, Bolboceratidae, rather than a subfamily. ITIS[1] currently treats Bolboceratinae as a subfamily of Geotrupidae.


These eight genera belong to the subfamily Bolboceratinae:[3]

Bolbelasmus Boucomont, 1911
Bolbocerastes Cartwright, 1953
Bolbocerosoma Schaeffer, 1906
Bolborhombus Cartwright, 1953
Bradycinetulus Cockerell, 1906
Eucanthus Westwood, 1848
Neoathyreus Howden and Martínez, 1963
Odonteus Samouelle, 1819


"Bolboceratinae Report". Integrated Taxonomic Information System. Retrieved 2019-05-08.
"Bolboceratinae subfamily Information". Retrieved 2019-05-08.
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
"Catalogue of Life". Retrieved 2018-03-28.

"GBIF". Retrieved 2018-03-28.

Further reading

Jameson, M.L. (2002). "Family 29: Geotrupidae". In Arnett, Ross H. Jr.; Thomas, Michael C.; Skelley, Paul E.; Frank, J. Howard (eds.). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. pp. 23–27. ISBN 978-0-8493-0954-0.
Ratcliffe, Brett; Jameson, Mary Liz. "Generic Guide to New World Scarab Beetles". Archived from the original on 2019-03-30. Retrieved 2019-07-02.
Lobl, I.; Smetana, A., eds. (2006). Catalogue of Palaearctic Coleoptera, Volume 3: Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books. ISBN 978-90-04-30914-2.
LeConte, J. L. (1861). Classification of the Coleoptera of North America. Smithsonian Miscellaneous Collections. Vol. 3. Smithsonian Institution. doi:10.5962/bhl.title.38459. ISBN 978-0665100550.

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