Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga
Familia: Carabidae
Subfamilia: Harpalinae
Supertribus: Pterostichitae
Tribus: Pterostichini
Genus: Pterostichus
Subgenera: P. (Abacidus) – P. (Adelosia) – P. (Aphaonus) – P. (Argutor) – P. (Bothriopterus) – P. (Circinatus) – P. (Cryobius) – P. (Cylindrocharis) – P. (Eosteropus) – P. (Euferonia) – P. (Feronina) – P. (Feroperis) – P. (Gastrosticta) – P. (Gutta) – P. (Huaius) – P. (Hypherpes) – P. (Koreonialoe) – P. (Lamenius) – P. (Lenapterus) – P. (Melanius) – P. (Metallophilus) – P. (Monoferonia) – P. (Morphnosoma) – P. (Morphocarabus) – P. (Morphohaptoderus) – P. (Nialoe) – P. (Orsonjohnsonus) – P. (Paraferonia) – P. (Petrophilus) – P. (Phonias) – P. (Plectes) – P. (Pseudethira) – P. (Pseudoferonina) – P. (Pseudomaseus) – P. (Pterostichus) – P. (Steropus) – P. (Tausternus) – P. (Tundraphilus) – P. (Vietosteropus)
Check: Leptoferonia
Overview of species
A
P. aanistschenkoi – P. aapsorum – P. abaciformis – P. abadiei – P. abagonensis – P. abasarukini – P. abasinus – P. abaxoides – P. abensis – P. abishirensis – P. abnormis – P. abrogatus – P. abudarini – P. acrogonus – P. acuspina – P. acutidens – P. acutipes – P. adatarasanus – P. adelaidae – P. ademidovae – P. adoxus – P. adstrictus – P. aedeagalis – P. aemiliae – P. aeneocupreus – P. aenescens – P. aeneus – P. aereipennis – P. aethiops – P. ageneralovi – P. agonus – P. aibgensis – P. aimaki – P. ainus – P. akaishicus – P. akkusianus – P. akitai – P. akozlovi – P. akozyrevi – P. aksekianus – P. alacer – P. albericus – P. alberti – P. alexandrovi – P. alexeji – P. algidus – P. altaianus – P. altaicus – P. altaiensis – P. alzonai – P. amadeus – P. amagisanus – P. amanoi – P. amaroides – P. ambigenus – P. ambiguus – P. amblypterus – P. amethystinus – P. amoenus – P. amorei – P. amurensis – P. analis – P. anatolicus – P. andorranus – P. andorranus – P. andreae – P. andreinii – P. andrewesi – P. angelieri – P. angladensis – P. angoarnigi – P. anguleus – P. angustistylis – P. angustus – P. annosus – P. anomostriatus – P. antepunctatus – P. anthracinus – P. antiquus – P. apenninus – P. apfelbecki – P. apiculatiphallus – P. aplutenkoi – P. aragonensis – P. aragonicus – P. aralarensis – P. araraticus – P. arator – P. arcanoides – P. arcanus – P. archangelicus – P. arcticola – P. arcuaticarinatus – P. arcuatus – P. argutoriformis – P. aristochroides – P. armenus – P. arnoldii – P. arribasi – P. arrowi – P. arrowianus – P. arsenjevi – P. arvernus – P. asahinus – P. asiaticus – P. askhicus – P. asturicus – P. asturicus – P. astutus – P. asumarokovi – P. asymmetricus – P. aterrimus – P. atratus – P. atrox – P. aubryi – P. audax – P. auratus – P. auriga – P. ausonicus – P. avadcharicus – P. avaricus – P. averenskii
B
P. baderlei – P. baenningeri – P. balachowskyi – P. baldensis – P. baldwini – P. balikliensis – P. balthasari – P. balu – P. bandotaro – P. bannaticus – P. barbarae – P. barbarinus – P. barbatus – P. barlensis – P. barri – P. barryorum – P. barthei – P. basilobatus – P. basipunctatus – P. batesi – P. battonii – P. baudii – P. belizini – P. bellatrix – P. belousovi – P. beneshi – P. berezowskii – P. bermamyticus – P. bernardi – P. bessoni – P. besucheti – P. beyeri – P. bhadravati – P. bhunetansis – P. bhutanensis – P. bicolor – P. bielzii – P. biexcisus – P. bifidiphallus – P. bifoveolatus – P. bigerricus – P. biimpressus – P. binotatus – P. biocryus – P. bischoffianus – P. bisetosus – P. bisulcitarsis – P. bituberculatus – P. blandulus – P. blodgettensis – P. blumenthali – P. bogatshevi – P. boisgiraudi – P. borcka – P. bosnicus – P. bowanus – P. brachylobus – P. brancuccii – P. breuili – P. brevicornis – P. brevilama – P. brevipennis – P. brevis – P. brevistylis – P. breviusculus – P. brittoni – P. bruckii – P. brunneipennis – P. bryanti – P. bryantoides – P. buglaniensis – P. bulgaricus – P. bungei – P. bureli – P. burmeisteri – P. bythiniensis
C
P. cabidochei – P. cabrerensis – P. cacumensis – P. californicus – P. caligans – P. calvitarsis – P. campania – P. canallatus – P. cancellatus – P. caniguensis – P. cantaber – P. cantabricus – P. cantalicus – P. capitatus – P. capito – P. capitolinus – P. cardioderus – P. caribou – P. carinatus – P. carinthiacus – P. carniolicus – P. carolinus – P. carpathicus – P. carradei – P. carri – P. carsticus – P. carthusianus – P. casalei – P. casaleianus – P. caspius – P. castaneus – P. castanipes – P. castor – P. catalonicus – P. catei – P. cathaicus – P. caudicalis – P. caucasicola – P. caucasicus – P. cavazzutii – P. cecchiniae – P. celtibericus – P. cervenkai – P. chainapaani – P. chameleon – P. champenoisi – P. championi – P. chasautianus – P. chechcirensis – P. chefsuricus – P. chenpengi – P. chinkiangensis – P. chipewyan – P. chogyesanus – P. chokaisanus – P. chotanensis – P. christinae – P. chujoi – P. chujoiellus – P. chungkingi – P. chydaeus – P. circulosus – P. clepsydra – P. cochlearis – P. cognatus – P. coiffaiti – P. colasi – P. colchicus – P. collaris – P. colonus – P. comatus – P. cometes – P. commixtiformis – P. commutabilis – P. comorus – P. compressus – P. comptorius – P. conaensis – P. confucius – P. congestus – P. consanguineus – P. consors – P. convexus – P. cordaticollis – P. cordatissimus – P. cordatus – P. cordifer – P. coreicus – P. corrusculus – P. coruscus – P. corvinus – P. costatus – P. crassiapex – P. crassiusculus – P. craterensis – P. cratocephalus – P. crebrepunctatus – P. crenicollis – P. crenulatopunctatus – P. creper – P. cribratus – P. cristatoides – P. cristatus – P. cristicaudis – P. cryobioides – P. cucujinus – P. cunibakus – P. cursor – P. curtatus – P. curvatus – P. cylindricus – P. cylindriformis
D
P. daghensis – P. daghestanus – P. daihizanus – P. daisenicus – P. dandongensis – P. dandonis – P. darvazicus – P. datshenkoae – P. dauricus – P. davidi – P. decastriensis – P. deceptrix – P. defossus – P. deino – P. delaroueriei – P. delicatus – P. dentatus – P. dentellus – P. denticaudis – P. depilatus – P. depressidorsis – P. depressiusculus – P. depressus – P. destructus – P. deuvei – P. deuvesianus – P. devillei – P. dhaulagiricus – P. dhorpatanicus – P. diana – P. dilatatus – P. diligendus – P. diligens – P. dilutipes – P. dipojranus – P. discrepans – P. dissimilis – P. distinctissimus – P. distinguendus – P. diversus – P. doderoi – P. dolens – P. doris – P. dorjulensis – P. dormitans – P. dostali – P. drescheri – P. drescoi – P. dudkoi – P. dufourii – P. dulcis – P. dundai – P. duratii – P. dux
E
P. ebeninus – P. ebenus – P. eboshiyamanus – P. ecarinatus – P. ehlersi – P. elmbergi – P. elongatus – P. elytralis – P. emei – P. empetricola – P. enasanus – P. enodis – P. enyo – P. eobius – P. eoyoritomus – P. epiroticus – P. erillensis – P. eriwanicus – P. esakii – P. escalevae – P. eschscholtzii – P. espanoli – P. estarragnensis – P. esuriens – P. ethnicus – P. euxinus – P. exceptus – P. exilis – P. eximius – P. exochus – P. expectatus – P. expeditus – P. externepunctatus – P. extraneus
F
P. falcispinus – P. faldermanni – P. fallettii – P. falli – P. farkaci – P. fasciatopunctatus – P. faunus – P. femoralis – P. femoratus – P. fencli – P. fenestratus – P. fenestrellanus – P. fenyesi – P. ferghanicus – P. fernquisti – P. ferreri – P. ferreroi – P. filipjevi – P. filum – P. findelii – P. firmus – P. flavofemoratus – P. focarilei – P. fornicatus – P. foveolatus – P. fractus – P. franzi – P. freyi – P. frigoris – P. fritzhiekei – P. fuchsi – P. fujimurai – P. fujisanus – P. fujitai – P. fukube – P. fuligineus – P. fulvescens – P. funakoshii – P. funerarius – P. funestes – P. fuscicornis
G
P. gagates – P. gagatinus – P. galae – P. galaecianus – P. galinae – P. gallega – P. gallopavo – P. ganesh – P. ganja – P. gassanus – P. gaudini – P. geberti – P. geojensis – P. georgi – P. gerdi – P. gerstlensis – P. ghilarovi – P. ghilianii – P. giacomazzoi – P. gibbicollis – P. giordanii – P. giraudi – P. giresuni – P. glabricollis – P. glabripennis – P. glacialis – P. glaferi – P. glasunowi – P. gliscans – P. globosus – P. glukhomanka – P. gobettii – P. goekensis – P. gompanus – P. gongga – P. goriensis – P. goschi – P. gotoensis – P. gracilior – P. gracilis – P. grajus – P. gravis – P. gschwendtneri – P. guedeli – P. guizhouensis – P. gujoensis – P. gwervaenus – P. gyrosus
H
P. haesitatus – P. haftorni – P. hagenbachii – P. hakusanus – P. haptoderoides – P. harbinensis – P. harmandi – P. harponifer – P. hartmanni – P. hatchi – P. hedkeinekus – P. heilongjiangensis – P. heinzi – P. heinzianus – P. hejkali – P. heptapotamicus – P. herculaneus – P. herzi – P. hespericus – P. hidanus – P. higonis – P. hikosanus – P. himifuriho – P. hirobane – P. hiramatsui – P. hirasawai – P. hisamatsui – P. hoelli – P. hoffmanni – P. homalonotus – P. honnoratii – P. honshuensis – P. hoplites – P. horioi – P. hornii – P. horstweiperti – P. horvatovichi – P. hozumii – P. huashanus – P. hubeicus – P. hudsonicus – P. huguai – P. humilis – P. hummeli – P. hungaricus – P. hustacheianus – P. hypogeus
I
P. iberlovi – P. idahoae – P. ienishteai – P. ignavus – P. ikizderensis – P. ikukoae – P. ikutanii – P. illigeri – P. illustris – P. imasakai – P. imitator – P. imitatus – P. immarginatus – P. impressicollis – P. impressus – P. imurai – P. inanis – P. incommodus – P. inexpectus – P. infaustus – P. infernalis – P. infimus – P. inflaticeps – P. ingusha – P. innshanensis – P. inopinatus – P. inopinus – P. insidatrix – P. insidiosus – P. insignicollis – P. insumiensis – P. intermedius – P. interruptus – P. intricatus – P. iranianus – P. irideus – P. iripennis – P. isabellae – P. ishidai – P. ishiharai – P. ishii – P. ishikawai – P. ishikawaioides – P. ishizuchiensis – P. ishizukai – P. isolatus – P. isumiensis – P. italicus – P. itoi – P. iwakiensis – P. iwasakii – P. iwawakianus – P. ixion
J
P. jacobinus – P. jacobsoni – P. jaechi – P. jaechianus – P. jakobsonianus – P. jaljaleensis – P. janatai – P. janbritoi – P. jankowskyi – P. jani – P. janoi – P. japanensis – P. japonicus – P. jarkendis – P. jeanneli – P. jelepus – P. jenisseicus – P. jiricola – P. jogaesanensis – P. johnsoni – P. josephi – P. juga – P. jugicola – P. jugivagus – P. jumlanus – P. jungens – P. jureceki – P. jurecekianus – P. jurinei – P. justusii
K
P. kacheticus – P. kadjaranze – P. kadleci – P. kadoudali – P. kajimurai – P. kajmakcalensis – P. kalabi – P. kalhys – P. kalo – P. kambaiti – P. kamtschaticus – P. kangchenjunga – P. kaniei – P. kaninensis – P. kantous – P. kapanovi – P. karasawai – P. karateghinicus – P. kassbergensis – P. kataevi – P. katashinensis – P. katoi – P. kawanoi – P. kaynashensis – P. keltiekus – P. kerzhneri – P. kiangsu – P. kiiensis – P. kimurai – P. kiritshenkoi – P. kirschenblatti – P. kitakamisanus – P. klapperichi – P. kleinfeldi – P. kleinfeldianus – P. kluchoriensis – P. koenigi – P. koenigianus – P. koheii – P. kokeilii – P. kolosovi – P. kolymensis – P. komalus – P. komiyai – P. kongosanus – P. kopetzi – P. korgei – P. korolkowi – P. kosakai – P. kotzebuei – P. kourili – P. krali – P. krasnobaevi – P. krasnopolensis – P. krestovyanus – P. krulikovskyi – P. kubanicus – P. kucerai – P. kultianus – P. kulzeri – P. kuporosovi – P. kurentzovi – P. kurikomasanus – P. kurnakovi – P. kurosai – P. kurosawai – P. kurosonis – P. kuwolsanensis – P. kvirensis – P. kyushuensis
L
P. laborans – P. labzuki – P. lacertus – P. lachrymosus – P. lacunosus – P. laevigatus – P. laevipunctatus – P. laferi – P. lama – P. lamuticus – P. lanista – P. larisae – P. larsi – P. lasserrei – P. lassulus – P. latemarginatus – P. laticollis – P. laticollis – P. latifianus – P. latistylis – P. latitemporis – P. latiusculus – P. latoricaensis – P. lattini – P. ledenikensis – P. leica – P. leleupi – P. leonianus – P. leonisi – P. leptis – P. lesourdi – P. lesticoides – P. letensis – P. levadensis – P. leviculus – P. lianhuaensis – P. licenti – P. liciniformis – P. lidara – P. lidarus – P. liguricus – P. liliputanus – P. lineatopunctatus – P. lingshanus – P. liodactylus – P. litae – P. liupanensis – P. ljubetensis – P. lobatus – P. lodosi – P. lombardus – P. lompokharianus – P. longinquus – P. longipennis – P. longipes – P. longiusculus – P. lubricus – P. lucidus – P. luctuosus – P. ludovici – P. lugi – P. luigionii – P. lukjanovitshi – P. lumensis – P. lustrans – P. lutschniki – P. lutshnikianus – P. lyroderus
M
P. macedonicus – P. macer – P. machulkai – P. macrocephalus – P. macrogenys – P. macrothorax – P. maderi – P. madidus – P. maeklini – P. magoides – P. magus – P. maichensis – P. mainardii – P. makalu – P. makolskii – P. malaisei – P. malissorum – P. malkini – P. mancus – P. mandibularoides – P. mandzhuricus – P. mannerheimii – P. marani – P. marginatus – P. marginellus – P. mariae – P. marinensis – P. martensi – P. maryseae – P. masahiroi – P. masatakai – P. mascarauxi – P. mascarauxi – P. masidai – P. masumotoi – P. matchai – P. mateui – P. matsumurai – P. mattolensis – P. matveichuki – P. maurusiacus – P. maximus – P. mayeti – P. mediocris – P. medvedevi – P. megacephalus – P. megaloderus – P. meisteri – P. melanarius – P. melanodes – P. melanoscelis – P. melas – P. mellyi – P. mendizabali – P. menetriesii – P. mercedianus – P. meridionalis – P. merkli – P. meskheticus – P. metallifer – P. mewakholensis – P. meyeri – P. micans – P. michailovi – P. michinoku – P. microbus – P. microcephalus – P. microphthalmus – P. micropoides – P. microps – P. middendorffi – P. miehei – P. migliaccioi – P. militaris – P. minax – P. ming – P. minor – P. minutulus – P. mirabilis – P. mirificus – P. miroshnikovi – P. miroslavi – P. mirus – P. miscellus – P. mitoyamanus – P. miyamai – P. mizunoyai – P. modicellus – P. moestus – P. molopsoides – P. mongolicus – P. mongoloaltaicus – P. monostigma – P. montadai – P. montanus – P. monticoloides – P. montigena – P. montisdeorum – P. montsenicus – P. morawitzianus – P. morio – P. morionides – P. mosaicus – P. mucronatus – P. muehlfeldii – P. muelleri – P. muellermotzfeldi – P. mujahedeeni – P. mukdenensis – P. mulensis – P. multinodosus – P. multipunctatus – P. multiseta – P. mundatus – P. mundus – P. musashiensis – P. mutoides – P. mutus
N
P. nadari – P. naganoensis – P. nakanei – P. namrum – P. naokii – P. napaea – P. nasuensis – P. nasui – P. necessarius – P. nechaevi – P. neglectus – P. negligens – P. neilgaimani – P. nemoralis – P. neobrunneus – P. nepalensis – P. nevadensis – P. nicaeensis – P. nigellatus – P. nigellus – P. niger – P. nigerrimus – P. nigripalpis – P. nigrita – P. nigrocaeruleus – P. nikodymi – P. nimbatidius – P. nishidai – P. nishiyamai – P. nivalis – P. nivicola – P. noburai – P. nodicornis – P. noguchii – P. nordqvisti – P. noricus – P. noui – P. novotnyorum – P. novus – P. nowitzkii
O
P. obesulus – P. obirensis – P. oblongiusculus – P. oblongoparallelus – P. oblongopunctatus – P. obsidianus – P. obtusus – P. occidentalis – P. occultus – P. ochoticus – P. ochraceus – P. ochsi – P. odontocnemis – P. odvarkai – P. oenotrius – P. ogaensis – P. ohbayashii – P. ohdaisanus – P. ohkawai – P. ohkurai – P. ohsawacavus – P. okiensis – P. okutamae – P. olafi – P. olegi – P. omogoensis – P. ompoensis – P. opacipennis – P. ophryoderus – P. ordinarius – P. ordinatoides – P. ordinatus – P. orduensis – P. oregonus – P. oreophilus – P. orestes – P. orientalis – P. orion – P. orionis – P. orman – P. orobius – P. osseticus – P. osseticus – P. ottomanus – P. ovalipennis – P. ovaliphallus – P. ovoideus
P
P. pachinus – P. palekhai – P. palgongsanus – P. palmi – P. panticulatus – P. pantomus – P. panzeri – P. paralleloides – P. parasimilis – P. parens – P. parkwon – P. parnassius – P. parviceps – P. parvicollis – P. pasanauricus – P. patris – P. patruelis – P. paulini – P. paulinoi – P. pavani – P. pecoudi – P. pecoudianus – P. pedemontanus – P. peilingi – P. pellegrinii – P. pemphredo – P. peninsularis – P. pensylvanicus – P. pentheri – P. percontator – P. pergracilis – P. perhoplites – P. perisi – P. perlamatus – P. perlidaghensis – P. perlutus – P. pertinax – P. petulans – P. pfizenmayeri – P. phaeopus – P. phaeus – P. piceolus – P. picipes – P. pilifer – P. piliferus – P. pilosus – P. pinguedineus – P. piniphilus – P. pithyusicus – P. planaticollis – P. planctus – P. planicola – P. planipennis – P. planiusculus – P. planus – P. platyderus – P. platyops – P. platypterus – P. plesiomorphus – P. poecilma – P. pohnerti – P. polygenus – P. ponticus – P. poppiusi – P. poppiusianus – P. porcellus – P. potanini – P. praedo – P. pratti – P. prevostii – P. primus – P. probus – P. procax – P. procephalus – P. procerulus – P. prolongatus – P. properans – P. protensiformis – P. protractus – P. provincialis – P. przewalskyi – P. pseudoastutus – P. pseudobarbarae – P. pseudodiversus – P. pseudodolens – P. pseudoharmandi – P. pseudojugivagus – P. pseudopachinus – P. pseudopedius – P. pseudopercus – P. pseudopilifer – P. pseudoplatyderus – P. pseudoplatysma – P. pseudorotundus – P. pseudosinensis – P. pseudostuxbergi – P. pueli – P. pulchellus – P. pulcher – P. pumilio – P. pumilus – P. pumpellyi – P. punctatus – P. punctibasalis – P. punctiger – P. punctisternus – P. punctiventris – P. pusillus – P. putus
Q
P. quadraticollis – P. quadrifoveolatus – P. quadriimpressus – P. quelpartensis – P. quezeli
R
P. radohanus – P. raptor – P. raraensis – P. rasilis – P. regeli – P. reiseri – P. reissi – P. relictus – P. rengensis – P. restrictus – P. reuteri – P. rhaeticus – P. rhanis – P. rhilensis – P. rifensis – P. riffensis – P. riparius – P. ripensis – P. robustistylis – P. rolex – P. ronchettii – P. rothi – P. rotundangulus – P. rotundithorax – P. rotundus – P. roubali – P. roubalianus – P. rousi – P. rousianus – P. rousselli – P. rubripalpis – P. rudestriatus – P. ruffoi – P. rufipes – P. rufitarsis – P. rufonitens – P. rufopiceus – P. rugibasis – P. rugiceps – P. rugosiceps – P. rugosipennis – P. rugosus – P. rutenus – P. rutilans – P. ryomoensis
S
P. sacheri – P. sachtlebeni – P. safonovae – P. sagarmatha – P. sagittalis – P. sakagamii – P. sakagutii – P. sakuraii – P. samniticus – P. santostamangi – P. sasajii – P. satoi – P. satsumanus – P. satunini – P. satyrus – P. saueri – P. saxicola – P. sayanus – P. scalptus – P. schaschli – P. schmidtii – P. schneideri – P. schodaicus – P. schoenherri – P. schoenmanni – P. schrettenbrunneri – P. schuelkei – P. schuleri – P. scitus – P. scurra – P. scutellaris – P. scuticollis – P. sediorcus – P. sejunctus – P. sejungendus – P. sellae – P. selmanni – P. semenowi – P. semirugosus – P. septemtrionalis – P. septentrionis – P. serbicus – P. seriatus – P. serripes – P. seticeps – P. setipes – P. setosicus – P. setosus – P. seunglaki – P. seungmoi – P. shaovuensis – P. sharanus – P. shavrini – P. shennongjianus – P. shibatai – P. shiibanus – P. shikatai – P. shilenkovi – P. shimizui – P. shinbodakensis – P. shingarevi – P. shirakamisan – P. shirakamisanus – P. shirozui – P. shojii – P. shotaroi – P. sierranus – P. silvestris – P. similatus – P. similis – P. simillimus – P. sincerus – P. sinensis – P. singularis – P. sinicus – P. sintanus – P. sinuatus – P. skoupyi – P. smaragdinus – P. sodalicius – P. sojaki – P. solskyi – P. songoricus – P. soperi – P. sotkaensis – P. sousae – P. sphodriformis – P. sphodrinus – P. spiculifer – P. spinolae – P. sponsor – P. spornyi – P. spraguei – P. ssemenovi – P. stagnatilis – P. stantonensis – P. stapedius – P. starcki – P. starckianus – P. staveni – P. sterbai – P. stictopleurus – P. stoeckleini – P. stomoides – P. straneellus – P. straneoi – P. strasseri – P. strenuus – P. stricticollis – P. strigipennis – P. strigosus – P. strupii – P. stygicus – P. styranus – P. styx – P. subabacipennis – P. subacutus – P. subaeneus – P. subgibbus – P. subiasi – P. subitus – P. sublaevis – P. sublucidus – P. subovatus – P. subparallelus – P. subrugosus – P. subsinuatus – P. subtilissimus – P. sudai – P. sudeticus – P. sudrei – P. suffusus – P. sugimurai – P. sulcitarsis – P. sumondakensis – P. sundukovi – P. sungariensis – P. superciliosus – P. surcoufi – P. suruganus – P. sutschanensis – P. svanicus – P. swaneticus – P. syleus – P. sylviae – P. symmetricus – P. szekessyianus – P. szetschuanensis
T
P. tachongi – P. taebaegsanus – P. tahirai – P. takadateyamanus – P. takahashii – P. takaosanus – P. taksangensis – P. taksonyis – P. talibani – P. tamarae – P. tamsii – P. tanakai – P. tanakaorum – P. tangkosis – P. tanjae – P. tantillus – P. taoi – P. taradakensis – P. tareumiut – P. tarsalis – P. tateishiyamanus – P. tatianae – P. tatricus – P. telavensis – P. temperei – P. tenenbaumianus – P. tenuimarginatus – P. tenuis – P. teretis – P. testaceus – P. tetralobatus – P. thanglaensis – P. theeli – P. thorectes – P. thorectoides – P. thunbergi – P. tichomirovi – P. tienmushanus – P. tiliaceoradix – P. todai – P. togyusanus – P. tokejii – P. tokmakovae – P. tokui – P. toledanoi – P. tomensis – P. tonkinensis – P. tosanus – P. tottoriensis – P. toyodai – P. transversalis – P. triangularis – P. trinarius – P. trinitensis – P. triseriatus – P. tristis – P. troglophilus – P. tropicalis – P. trubilini – P. truncatus – P. truncatus – P. tscherkessicus – P. tschetschenicus – P. tschitscherinianus – P. tsukubasanus – P. tsurugaensis – P. tsurugiyamanus – P. tuberculifer – P. tuberculiger – P. tuberculofemoratus – P. tuberifer – P. tuberosus – P. tukchensis – P. tulgoeti – P. tulobalu – P. tumescens – P. tundrae – P. turanensis – P. turcicus – P. turcomannicus
U
P. uchiyamai – P. uedaorum – P. uenoi – P. ulubeyensis – P. umbralis – P. umbricola – P. unctulatus – P. uralensis – P. urengaicus – P. ussuriensis
V
P. vagepunctatus – P. validiceps – P. validior – P. validiusculus – P. validus – P. vallierensis – P. vandykei – P. vanvolxemi – P. variabilis – P. variicornis – P. variolatus – P. variseta – P. vasconicus – P. vecors – P. ventralis – P. ventricosus – P. vermiculosus – P. vernalis – P. vicinus – P. virescens – P. viridellus – P. vitosensis – P. vladivostokensis – P. vseteckai
W
P. walcotti – P. walteri – P. watanabei – P. weigeli – P. weiperti – P. wellensii – P. wittmeri – P. woodi – P. woongbii – P. woronowi – P. wrasei
X
P. xatartii
Y
P. yahikosanus – P. yakushimanus – P. yamajii – P. yamauchii – P. yamizosanus – P. yanoi – P. yatsuensis – P. ybousqueti – P. yokohamae – P. yorikoae – P. yoritomus – P. yosemitensis – P. yoshidai – P. yoshikawai – P. yoshizawai – P. yulongshanensis – P. yunnanensis – P. yvanii
Z
P. zamotajlovi – P. zhejiangensis – P. ziegleri – P. zoiai – P. zolotarewi – P. zyzzovi
Note:
Add to (subgenus Circinatus): P. adelphus – P. ailaoicus – P. camelus – P. cavazzutianus
(new replacement name) – P. c. mianningensis – P. dimorphus – P. maitreya – P. miao – P. tumulus – P. wangjiani – P. yan – P. yuxiaodongi – P. zhygealu
Name
Pterostichus Bonelli, 1810: Tabula Synoptica
Type species: Carabus fasciatopunctatus Creutzer, 1799 (designated by Curtis 1828: plate 196)
References
Primary references
Bonelli, F.A. 1810. Observations Entomologiques, Première partie. Mémoires de l’Academie des Sciences de Turin 18: 21–78, Tabula Synoptica. BHL Reference page.
Additional references
Allegro, G. & Sciaky, R. 2010. Review of the Chinese subgenus Circinatus of Pterostichus, with description of six new species (Coleoptera, Carabidae). Bollettino del Museo Civico di Storia Naturale di Verona 34: 3–18. Full article (PDF).
Budilov, P.V. 2022. A new species of the genus Pterostichus Bonelli, 1810 (Coleoptera: Carabidae) from the Russian Far East. Far Eastern Entomologist 446: 24-28. DOI: 10.25221/fee.446.4. Open access. Full article (PDF). Reference page.
Chaladze, G., Kalatozishvili, L., Janiashvili, Z. & Bakuradze, G. 2017. Pterostichus neilgaimani sp. nov., a new species of ground beetles (Coleoptera: Carabidae) from relict sacred grove in Eastern Georgia. Zootaxa 4329(1): 91–95. DOI: 10.11646/zootaxa.4329.1.6 Full article (PDF). Reference page.
Dorjderem, S., Shi, H-L. & Liang, H-B. 2020. The genus Pterostichus in China III: a brief review of subgenus Chinapterus Berlov (Coleoptera, Carabidae) with descriptions of two new species. ZooKeys, 953: 61–83. DOI: 10.3897/zookeys.953.52282 Open access Reference page.
Fedorenko, D.N. 2020. New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam. Russian Entomological Journal 29(1): 38-52. DOI: 10.15298/rusentj.29.1.06. Open access. Reference page.
Kabak, I.I. 2021. New species of the genus Pterostichus Bonelli, 1810 (Coleoptera: Carabidae) from North-Western China. Caucasian Entomological Bulletin 17(2): 307–313. Full article (PDF). Reference page.
LaBonte, J.R. 2013. Pterostichus (Atilloferonia) diana LaBonte (Coleoptera: Carabidae: Pterostichini), a replacement name for P. (A.) lanei (Hatch, 1935), and validity and redescription of P. (A.) malkini (Hatch, 1953). Zootaxa 3682(4): 563–571. DOI: 10.11646/zootaxa.3682.4.7 Rreference page.
Morita, S., Ohkawa, H. & Kurihara, T. 2013. Two New Macrocephalic Pterostichine Carabids (Coleoptera, Carabidae) from Central Japan. Elytra (n.s.), 3(1): 9-17. Full article (PDF). Reference page.
Sasakawa, K. & Itô, H. 2017. Two new species of the Pterostichus macrogenys species group (Coleoptera, Carabidae) discovered in shallow subterranean habitats in northern Honshu, Japan. Subterranean Biology, 21: 47–56. DOI: 10.3897/subtbiol.21.11155 Open access Reference page.
Morita, S. & Ohkawa, H. 2010. A new Pterostichus (Coleoptera, Carabidae) from Gifu Prefecture, Central Japan. Elytra 38(1): 99–103. [not seen]
Shi, H.-L., Sciaky, R., Liang, H. & Zhou, H.-Z. 2013. A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa 3664(2): 101–135. DOI: 10.11646/zootaxa.3664.2.1 Reference page.
Shi, H-L. & Liang, H-B. 2015. The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini). Zookeys 536: 1–92. DOI: 10.3897/zookeys.536.5982 Full article. Reference page.
Sun, X-J., Shi, H-L., Sang, W-G. & Axmacher, J.C. 2018. Two new species of Feroperis Lafer (Carabidae, Pterostichus) from China, with a key to all known Chinese species in this subgenus. Zookeys, 799: 95–114. DOI: 10.3897/zookeys.799.28834 Reference page.
Tian, M.Y., Huang, S.B., Chen, M.Z. & Ding, K.J. 2019. Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae). Annales de la Société entomologique de France (N.S.) 55(1): 1–16. DOI: 10.1080/00379271.2018.1546554. PaywallReference page.
Toda, N. 2012. Two new macrocephalic Pterostichines (Coleoptera, Carabidae) from Central Japan. Elytra (n.s.) 2(1): 119–125. PDF Reference page.
Wrase, D.W. & Schmidt, J. 2006. Description of a new subgenus and five new species of the genus Pterostichus Bonelli, 1810 (Coleoptera, Carabidae, Pterostichini) from Northern Myanmar and Yunnan. Veröffentlichung des Naturkundemuseums Erfurt 25: 203–215. [not seen]
Yin,W-Q., Shi, H-L. & Liang, H-B. 2021. Revision of the Chinese species of subgenus Koreonialoe (Coleoptera: Carabidae: Pterostichus), with descriptions of two new species. Zookeys 1063ː 1–21. DOI: 10.3897/zookeys.1063.69942 Open access Reference page.
Zamotajlov, A.S., Solodovnikov, I.A. & Fominykh, D.D. 2015. Description of two new species related to Pterostichus (Aphaonus) arcanoides Lorenz, 1998 from the West Transcaucasia (Coleoptera: Carabidae: Pterostichini). Zootaxa 3999(1): 111–124. DOI: 10.11646/zootaxa.3999.1.7 Preview (PDF). Reference page.
Links
Pterostichus – Taxon details on Biological Library (BioLib).
Pterostichus – Taxon details on Global Biodiversity Information Facility (GBIF).
Berlov, O. Ground beetles of the genus Pterostichus (Carabidae): atlas of beetles of Russia. Beetles (Coleoptera) and coleopterists.
Vernacular names
čeština: Střevlíček
Deutsch: Grabkäfer
suomi: Sysikiitäjäiset
русский: Птеростих
Pterostichus is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer slightly moist, sandy soil although preferences differ between species.[1]
Species
Pterostichus contains the following species:[2][3][4][5]
Pterostichus aanistschenkoi O. Berlov & E. Berlov, 1999
Pterostichus aapsorum Belousov, 1991
Pterostichus abaciformis Straneo, 1955
Pterostichus abagonensis Reitter, 1896
Pterostichus abasarukini O. Berlov & E. Berlov, 1996
Pterostichus abasinus Belousov, 1991
Pterostichus abaxoides Dejean, 1828
Pterostichus abensis Morita, 2010
Pterostichus abishirensis Belousov, 1991
Pterostichus abnormis (J.Sahlberg, 1880)
Pterostichus abrogatus Scudder, 1890
Pterostichus abudarini O. Berlov & E. Berlov, 1997
Pterostichus acrogonus (Chaudoir, 1843)
Pterostichus acuspina (Tschitscherine, 1901)
Pterostichus acutidens (Fairmaire, 1889)
Pterostichus acutipes Barr, 1971
Pterostichus adatarasanus Sasakawa, 2005
Pterostichus ademidovae O.Berlov, 1996
Pterostichus adoxus (Say, 1823)
Pterostichus adstrictus Eschscholtz, 1823
Pterostichus adulterinus Wrase & J.Schmidt, 2006
Pterostichus aemiliae Facchini & Sciaky, 2000
Pterostichus aeneocupreus (Fairmaire, 1887)
Pterostichus aenescens Chaudoir, 1850
Pterostichus aeneus Fischer von Waldheim, 1829
Pterostichus aereipennis (Solsky, 1871)
Pterostichus aethiops (Panzer, 1796)
Pterostichus agilis Allegro & Sciaky, 2010
Pterostichus agonus G.horn, 1880
Pterostichus aibgensis Starck, 1890
Pterostichus aimaki Jedlicka, 1968
Pterostichus ainus Plutenko, 2005
Pterostichus akitai Morita, 2004
Pterostichus akkusianus Kirschenhofer, 1981
Pterostichus akozyrevi O. Berlov & E. Berlov, 1999
Pterostichus aksekianus (Straneo, 1988)
Pterostichus alacer A.Morawitz, 1862
Pterostichus alamedae (Casey, 1918)
Pterostichus alexandrovi Lafer, 1979
Pterostichus alexeji Zamotajlov & Kryzhanovskij, 1992
Pterostichus algidus Leconte, 1852
Pterostichus altaianus Jedlicka, 1958
Pterostichus altaicus (Germar, 1824)
Pterostichus altaiensis Poppius, 1906
Pterostichus amadeus Bousquet, 2012
Pterostichus amagisanus Tanaka & Ishida, 1972
Pterostichus amanoi Nakane, 1968
Pterostichus amaroides Dejean, 1828
Pterostichus ambigenus Bates, 1883
Pterostichus ambiguus (Fairmaire, 1858)
Pterostichus amblypterus Chaudoir, 1868
Pterostichus amethystinus Mannerheim, 1843
Pterostichus amoenus Dejean, 1828
Pterostichus ampliatus Bates, 1882
Pterostichus amurensis Poppius, 1906
Pterostichus analis Jedlicka, 1963
Pterostichus anatolicus Jedlicka, 1963
Pterostichus andreae (Tschitscherine, 1897)
Pterostichus andreinii Dodero, 1922
Pterostichus andrewesi Jedlicka, 1931
Pterostichus angoarnigi Morvan, 1994
Pterostichus angustus (Dejean, 1828)
Pterostichus annamita (Straneo, 1939)
Pterostichus annosus Casey, 1913
Pterostichus anomalus Chaudoir, 1878
Pterostichus anomostriatus Sciaky, 1995
Pterostichus antepunctatus Straneo, 1949
Pterostichus anthracinus (Illiger, 1798)
Pterostichus apenninus Dejean, 1831
Pterostichus apfelbecki Csiki, 1908
Pterostichus apiculatiphallus Nemoto, 1988
Pterostichus aralarensis Espanol & Mateu, 1945
Pterostichus araraticus Kirschenhofer, 1987
Pterostichus arator (Faldermann, 1836)
Pterostichus arcanoides Lorens, 1998
Pterostichus arcanus Casey, 1913
Pterostichus archangaicus Shilenkov, 2000
Pterostichus arcticola (Chaudoir, 1868)
Pterostichus arcuaticarinatus Kasahara, 1986
Pterostichus argutoriformis Poppius, 1906
Pterostichus aristochroides Deuve, 2006
Pterostichus armenus (Faldermann, 1836)
Pterostichus arnoldii Erjiomin, 1996
Pterostichus arribasi Ortuño, 1991
Pterostichus arrowi Jedlicka, 1936
Pterostichus arrowianus Jedlicka, 1938
Pterostichus arsenjevi Lafer, 1979
Pterostichus arvernus Jeannel, 1937
Pterostichus asahinus Habu & Baba, 1960
Pterostichus asiaticus Poppius, 1906
Pterostichus asturicus (Jeanne, 1965)
Pterostichus astutus (Tschitscherine, 1903)
Pterostichus asymmetricus Bates, 1883
Pterostichus aterrimus (Herbst, 1784)
Pterostichus atratus Gistel, 1857
Pterostichus atrox (Andrewes, 1937)
Pterostichus aubryi Coiffait, 1952
Pterostichus audax (Tschitscherine, 1895)
Pterostichus auratus Heer, 1837
Pterostichus auriga Ball, 1962
Pterostichus avaricus Kryzhanovskij & Abdurakhmanov, 1983
Pterostichus aztecus Tschitscherine, 1897
Pterostichus baenningeri Jedlicka, 1931
Pterostichus balachowskyi Morvan, 1972
Pterostichus baldwini (Casey, 1924)
Pterostichus balikliensis Hovorka & Skoupy, 2007
Pterostichus balthasari Jedlicka, 1937
Pterostichus balu Schmidt, 2009
Pterostichus bandotaro Tanaka, 1958
Pterostichus bannaticus Dejean, 1828
Pterostichus barbarae Sciaky & Facchini, 2003
Pterostichus barbarinus Casey, 1913
Pterostichus barbatus Fischer von Waldheim, 1829
Pterostichus bargusinicus Shilenkov, 2000
Pterostichus barlensis Straneo, 1935
Pterostichus barri Bousquet, 2006
Pterostichus barryorum Ball, 1962
Pterostichus basilobatus Sasakawa, 2005
Pterostichus basipunctatus Straneo, 1955
Pterostichus batesi (Tschitscherine, 1894)
Pterostichus belizini Lutshnik, 1933
Pterostichus bellatrix (Tschitscherine, 1895)
Pterostichus belousovi Kryzhanovskij, 1989
Pterostichus beneshi Sciaky, 1996
Pterostichus berezowskii (Tschitscherine, 1898)
Pterostichus besucheti (Straneo, 1988)
Pterostichus beyeri Vandyke, 1925
Pterostichus bhadravati Jedlicka, 1965
Pterostichus bhunetansis Davies, 2004
Pterostichus bhutanensis Morvan, 1980
Pterostichus bicolor Fischer von Waldheim, 1829
Pterostichus bielzii (Fuss, 1858)
Pterostichus biexcisus Straneo, 1955
Pterostichus bifidiphallus Nemoto, 1988
Pterostichus bifoveolatus Park; Kwon & Lafer, 1996
Pterostichus binotatus Fischer von Waldheim, 1829
Pterostichus biocryus Ball, 1962
Pterostichus bischoffianus Jedlicka, 1936
Pterostichus bisetosus (Straneo, 1938)
Pterostichus bisulcitarsis N.Ito, 2012
Pterostichus bituberculatus Tschitscherine, 1899
Pterostichus blandulus L.Miller, 1859
Pterostichus blodgettensis Will, 2007
Pterostichus blumenthali Heinz, 1965
Pterostichus bogatshevi (Kryzhanovskij, 1983)
Pterostichus borcka Jedlicka, 1963
Pterostichus bousqueti Bergdahl, 2011
Pterostichus bowanus Sciaky, 1997
Pterostichus brachylobus Kavanaugh and LaBonte, 2006
Pterostichus brachypterus Chaudoir, 1878
Pterostichus brancuccii Straneo, 1982
Pterostichus breuili Jeannel, 1919
Pterostichus brevicornis Kirby, 1837
Pterostichus brevilama Straneo, 1989
Pterostichus brevipennis (Chevrolat, 1840)
Pterostichus brevis (Duftschmid, 1812)
Pterostichus breviusculus R.F. Sahlberg, 1844
Pterostichus brittoni Habu, 1958
Pterostichus bruckii Schaum, 1859
Pterostichus brunneipennis Straneo, 1955
Pterostichus bryanti (Vandyke, 1951)
Pterostichus bryantoides Ball, 1962
Pterostichus bucolicus Casey, 1913
Pterostichus buglaniensis Kirschenhofer, 1981
Pterostichus bullatus Allegro & Sciaky, 2010
Pterostichus bungei (Tschitscherine, 1894)
Pterostichus bureli Morvan, 1994
Pterostichus burjaticus Poppius, 1906
Pterostichus burkhan O.Berlov & Anistschenko, 2005
Pterostichus burmeisteri Heer, 1838
Pterostichus bythiniensis (Maran, 1944)
Pterostichus cacumensis Ball, 1966
Pterostichus californicus (Dejean, 1828)
Pterostichus caligans G.Horn, 1891
Pterostichus calvitarsis Breit, 1912
Pterostichus campania Andrewes, 1937
Pterostichus campbelli Bousquet, 1985
Pterostichus canallatus Casey, 1913
Pterostichus cancellatus (Motschulsky, 1860)
Pterostichus cantabricus Schaufuss, 1862
Pterostichus cantalicus (Chaudoir, 1868)
Pterostichus capitatus Chaudoir, 1850
Pterostichus capito (Tschitscherine, 1900)
Pterostichus capitolinus Kurnakov, 1962
Pterostichus caribou Ball, 1962
Pterostichus carinatus Sciaky & Facchini, 2003
Pterostichus carinthiacus Meschnigg, 1946
Pterostichus carolinus Darlington, 1932
Pterostichus carradei Gautier des Cottes, 1866
Pterostichus carri Angus; Galian; Wrase & Chaladze, 2009
Pterostichus carsticus Kasahara & Y.ito, 1989
Pterostichus casalei Straneo, 1984
Pterostichus casaleianus Kirschenhofer, 1991
Pterostichus caspius (Ménétriés, 1832)
Pterostichus castaneus (Dejean, 1828)
Pterostichus castanipes (Ménétriés, 1843)
Pterostichus castor Goulet and Bousquet, 1983
Pterostichus catalonicus J.Daniel, 1906
Pterostichus catei Sciaky & Wrase, 1997
Pterostichus cathaicus Sciaky, 1994
Pterostichus caucasicola (Tschitscherine, 1893)
Pterostichus caucasicus (Ménétriés, 1832)
Pterostichus caudicalis (Say, 1823)
Pterostichus cavazzutii Allegro & Sciaky, 2010
Pterostichus cavicollis Straneo, 1984
Pterostichus cecchiniae Jakobson, 1907
Pterostichus cervenkai Sciaky, 1994
Pterostichus chainapaani J.Schmidt, 2005
Pterostichus chameleon Motschulsky, 1866
Pterostichus champenoisi Croissandeau, 1893
Pterostichus championi Andrewes, 1926
Pterostichus chasautianus Vysoky, 1984
Pterostichus chechcirensis Lafer, 1979
Pterostichus chefsuricus Reitter, 1896
Pterostichus chenpengi Li, 1992
Pterostichus chinkiangensis Kirschenhofer, 1991
Pterostichus chipewyan Ball, 1962
Pterostichus chogyesanus Park; Kwon & Lafer, 1996
Pterostichus chokaisanus Sasakawa, 2009
Pterostichus chotanensis (Tschitscherine, 1893)
Pterostichus chujoi Habu, 1959
Pterostichus chujoiellus Jedlicka, 1962
Pterostichus chungkingi Jedlicka, 1932
Pterostichus chydaeus (Tschitscherine, 1897)
Pterostichus circulosus Lindroth, 1966
Pterostichus clepsydra Sciaky & Wrase, 1997
Pterostichus cochlearis Hacker, 1968
Pterostichus cognatus (Dejean, 1831)
Pterostichus coimbatoris Jedlicka, 1969
Pterostichus colasi Jeannel, 1937
Pterostichus colchicus Chaudoir, 1850
Pterostichus collaris Motschulsky, 1844
Pterostichus colonus (Bates, 1883
Pterostichus comatus Shi & Sciaky, 2013
Pterostichus cometes Andrewes, 1936
Pterostichus commixtiformis Roubal, 1920
Pterostichus commutabilis (Motschulsky, 1866)
Pterostichus comorus Jedlicka, 1932
Pterostichus compressus Rost, 1892
Pterostichus comptorius Gistel, 1857
Pterostichus conaensis J.Schmidt & Tian, 2011
Pterostichus confucius Sciaky & Wrase, 1997
Pterostichus congestus (Menetries, 1843)
Pterostichus consanguineus (Chaudoir, 1878)
Pterostichus consors (Tschitscherine, 1893)
Pterostichus convexus Gebler, 1847
Pterostichus coracinus (Newman, 1838)
Pterostichus cordaticollis Heyden, 1884
Pterostichus cordatissimus (Straneo, 1937)
Pterostichus cordifer Reitter, 1896
Pterostichus coreicus Jedlicka, 1962
Pterostichus corrusculus Leconte, 1873
Pterostichus coruscus Tschitscherine, 1895
Pterostichus corvinus (Dejean, 1828)
Pterostichus costatus (Ménétriés, 1851)
Pterostichus crassiapex Shi & Sciaky, 2013
Pterostichus crassiusculus (Chaudoir, 1868)
Pterostichus craterensis (Hatch, 1949)
Pterostichus cratocephalus Tschitscherine, 1897
Pterostichus crebrpunctatus Stranao, 1937
Pterostichus crenicollis Leconte, 1873
Pterostichus crenulatopunctatus (R.F.Sahlberg, 1844)
Pterostichus creper (Tschitscherine, 1902
Pterostichus cribratus (Dejean, 1828)
Pterostichus cristatoides Straneo, 1955
Pterostichus cristatus (L.Dufour, 1820)
Pterostichus cristicaudis Kurnakov, 1962
Pterostichus cryobioides Chaudoir, 1868
Pterostichus cubensis Darlington, 1937
Pterostichus cucujinus Reitter, 1892
Pterostichus cuneatulus Casey, 1913
Pterostichus cunibakus Davies, 2004
Pterostichus cursor Dejean, 1828
Pterostichus curtatus Fairmaire, 1886
Pterostichus curvatus Sciaky & Facchini, 2003
Pterostichus cylindricus (Herbst, 1784)
Pterostichus cylindriformis Reitter, 1887
Pterostichus daghestanus Reitter, 1896
Pterostichus daihizanus Ishida, 1968
Pterostichus daisenicus Ishida, 1958
Pterostichus dandongensis Kirschenhofer, 1997
Pterostichus dandonis Kasahara, 1989
Pterostichus darvasicus Kryzhanovskij, 1968
Pterostichus darvazicus Kryzhanovskij, 1968
Pterostichus datshenkoae Sundukov, 2013
Pterostichus dauricus (Gebler, 1832)
Pterostichus davidi Tschitscherine, 1897
Pterostichus davshensis Shilenkov, 2000
Pterostichus decastriensis Lafer, 1979
Pterostichus deceptrix Tschitscherine, 1898
Pterostichus defossus Bates, 1883
Pterostichus deino Will, 2007
Pterostichus dentellus Facchini & Sciaky, 2003
Pterostichus denticaudis Kurnakov, 1962
Pterostichus dentifer Allegro & Sciaky, 2010
Pterostichus depilatus (Bates, 1889)
Pterostichus depletus Scudder, 1900
Pterostichus depressidorsis Reitter, 1896
Pterostichus destitutus Scudder, 1890
Pterostichus destructus Scudder, 1890
Pterostichus deuvei Lassalle, 1985
Pterostichus deuvesianus Morvan, 1995
Pterostichus devillei Puel, 1924
Pterostichus dhorpatanicus Straneo, 1977
Pterostichus diabolus Casey, 1913
Pterostichus diana LaBonte, 2013
Pterostichus diligendus (Chaudoir, 1868)
Pterostichus diligens (Sturm, 1824)
Pterostichus dilutipes (Motschulsky, 1844)
Pterostichus dipojranus (Straneo, 1988)
Pterostichus discrepans A.Morawitz, 1862
Pterostichus dissimilis (A. & G.B.Villa, 1833)
Pterostichus distinctissimus Jedlicka, 1940
Pterostichus diversus (Fairmaire, 1886)
Pterostichus dolens (Tschitscherine, 1900)
Pterostichus doris Jedlicka, 1962
Pterostichus dorjulensis Morvan, 1978
Pterostichus dormitans Scudder, 1890
Pterostichus dostali Kirschenhofer, 1981
Pterostichus drescheri (Fischer, 1817)
Pterostichus drescoi Nègre, 1957
Pterostichus dudkoi Sundukov, 2013
Pterostichus dufourii (Dejean, 1828)
Pterostichus dulcis Bates, 1883
Pterostichus dundai Sciaky, 1994
Pterostichus duratii A. & G.B.Villa, 1835
Pterostichus dux L.Schaufuss, 1862
Pterostichus ebeninus (Dejean, 1828)
Pterostichus ebenus (Quensel, 1806)
Pterostichus eboshiyamanus Sasakawa, 2009
Pterostichus ecarinatus Hatch, 1936
Pterostichus ehlersi Heyden, 1881
Pterostichus elmbergi Poppius, 1908
Pterostichus elongatus (Duftschmid, 1812)
Pterostichus emei Sciaky, 1994
Pterostichus empetricola (Dejean, 1828)
Pterostichus enasanus Morita, 2007
Pterostichus enodis Bousquet, 1992
Pterostichus enyo Will, 2007
Pterostichus eobius (Tschitscherine, 1899)
Pterostichus eoyoritomus Sasakawa, 2009
Pterostichus epiroticus Csiki, 1930
Pterostichus eriwanicus Tschitscherine, 1897
Pterostichus esakii Ishida, 1959
Pterostichus eschscholtzii (Germar, 1824)
Pterostichus espanoli J. Vives & E. Vives, 1977
Pterostichus esuriens Casey, 1913
Pterostichus ethnicus Gistel, 1857
Pterostichus euxinus Straneo, 1935
Pterostichus evanescens Casey, 1918
Pterostichus exceptus J.R. Sahlberg, 1887
Pterostichus exilis Mateu, 2001
Pterostichus eximius A.Morawitz, 1862
Pterostichus exochus Andrewes, 1930
Pterostichus expeditus Tschitscherine, 1898
Pterostichus externepunctatus (Dejean, 1828)
Pterostichus falcispinus Sasakawa, 2005
Pterostichus faldermanni Schatzmayr, 1929
Pterostichus fallax (Dejean, 1828)
Pterostichus fallettii Sciaky, 1996
Pterostichus falli Vandyke, 1925
Pterostichus farkaci Sciaky, 1997
Pterostichus fasciatopunctatus (Creutzer, 1799)
Pterostichus faunus Kurnakov, 1962
Pterostichus femoralis (Kirby, 1837)
Pterostichus fencli Dvorak, 1995
Pterostichus fenestratus Kasahara & Ohtani, 1989
Pterostichus fenyesi Csiki, 1930
Pterostichus ferghanicus Kabak, 2001
Pterostichus fernquisti Wickham, 1931
Pterostichus ferreri (Español & Mateu, 1942)
Pterostichus ferreroi Straneo, 1989
Pterostichus filipjevi (Lutshnik, 1928)
Pterostichus filum Tschitscherine, 1897
Pterostichus findelii (Dejean, 1828)
Pterostichus firmus Lutshnik, 1933
Pterostichus flavofemoratus (Dejean, 1828)
Pterostichus focarilei Casale & Giachino, 1985
Pterostichus fornicatus Kolenati, 1845
Pterostichus foveolatus (Duftschmid, 1812)
Pterostichus fractus Scudder, 1890
Pterostichus franzi Nègre, 1955
Pterostichus fritzhiekei J.Schmidt, 1994
Pterostichus fuchsi Schaeffer, 1910
Pterostichus fujimurai Habu, 1958
Pterostichus fujisanus Tanaka & Suga, 1972
Pterostichus fujitai Tanaka & Ishida, 1972
Pterostichus fukube Sugimura, 2005
Pterostichus fuligineus A.Morawitz, 1862
Pterostichus fulvescens (Motschulsky, 1844)
Pterostichus funakoshii Morita, 2007
Pterostichus funestes Csiki, 1930
Pterostichus fuscicornis (Reiche & Saulcy, 1855)
Pterostichus gagates (Hope, 1831)
Pterostichus galae Farkac & Plutenko, 1996
Pterostichus galaecianus Lauffer, 1909
Pterostichus galinae Kabak, 1992
Pterostichus gallega (Fairmaire, 1859)
Pterostichus gallopavo Sciaky & Wrase, 1997
Pterostichus ganesh J.Schmidt, 2005
Pterostichus ganja J.Schmidt, 1995
Pterostichus gaoligongensis Wrase & J.Schmidt, 2006
Pterostichus gassanus Sasakawa, 2009
Pterostichus geberti Sciaky &, Wrase, 1997
Pterostichus geojensis Sasakawa; Kim & Kubota, 2006
Pterostichus georgi Straneo, 1963
Pterostichus gerdi Schmidt, 2009
Pterostichus gerstlensis Ball, 1962
Pterostichus ghilarovi Kryzhanovskij, 1988
Pterostichus ghilianii Putzeys, 1846
Pterostichus giacomazzai Sciaky, 1994
Pterostichus giacomazzoi Sciaky, 1994
Pterostichus gibbicollis (Motschulsky, 1844)
Pterostichus giresuni Jedlicka, 1965
Pterostichus glabricollis Jedlicka, 1962
Pterostichus glabripennis Jedlicka, 1962
Pterostichus glacialis Brisout de Barneville, 1863
Pterostichus glaferi O.Berlov & E.Berlov, 1996
Pterostichus gliscans Casey, 1913
Pterostichus glukhomanka Sundukov, 2013
Pterostichus gobettii Straneo, 1983
Pterostichus gompanus Straneo, 1983
Pterostichus gongga Sciaky, 1997
Pterostichus goriensis (Tschitscherine, 1897)
Pterostichus goschi Jedlicka, 1930
Pterostichus gotoensis Kasahara & Matsumoto, 1990
Pterostichus gracilior Leconte, 1873
Pterostichus gracilis (Dejean, 1828)
Pterostichus grajus (Dejean, 1828)
Pterostichus gravis Jedlicka, 1939
Pterostichus gregalis Casey, 1913
Pterostichus gromykoi Sundukov, 2005
Pterostichus guizhouensis Sciaky, 1997
Pterostichus gwervaenus Morvan, 1995
Pterostichus gyrosus (Motschulsky, 1866)
Pterostichus haesitatus Fairmaire, 1889
Pterostichus haftorni Lindroth, 1969
Pterostichus hagenbachii Sturm, 1824
Pterostichus hakusanus Kasahara, 1989
Pterostichus hamiltoni G. Horn, 1880
Pterostichus haptoderoides (Tschitscherine, 1889)
Pterostichus harmandi (Tschitscherine, 1900)
Pterostichus harponifer Tanaka, 1987
Pterostichus hartmanni J.Schmidt, 1995
Pterostichus hatchi Hacker, 1968
Pterostichus hedkeinekus Morvan, 1995
Pterostichus heilongjiangensis Li, 1992
Pterostichus heinzi Jedlicka, 1965
Pterostichus heinzianus Sciaky, 1996
Pterostichus hejkali B.Gueorguiev & Skoupy, 2011
Pterostichus heptapotamicus (Lutshnik, 1927)
Pterostichus herculaneus Mannerheim, 1843
Pterostichus herzi Poppius, 1906
Pterostichus hidanus Morita, 2010
Pterostichus hikosanus Kasahara, 1994
Pterostichus himifuriho Morita, 2007
Pterostichus hiramatsui Morita, 2010
Pterostichus hirasawai Morita, 2010
Pterostichus hirobane Habu & Baba, 1958
Pterostichus hisamatsui Ishida & Shibata, 1961
Pterostichus hoelli Schmidt, 2009
Pterostichus homalonotus Tschitscherine, 1894
Pterostichus honnoratii (Dejean, 1828)
Pterostichus hoplites (Bates, 1883)
Pterostichus hornii Leconte, 1873
Pterostichus horstweiperti J.Schmidt & Guorguiev, 2013
Pterostichus horvatovichi Kirschenhofer, 1991
Pterostichus hozumii Ishida, 1961
Pterostichus huashanus Sciaky, 1994
Pterostichus hubeicus Facchini & Sciaky, 2003
Pterostichus hudsonicus Leconte, 1863
Pterostichus humboldti Casey, 1913
Pterostichus humidulus (Vandyke, 1943)
Pterostichus humilis Casey, 1913
Pterostichus hummeli Jedlicka, 1935
Pterostichus hungaricus (Dejean, 1828)
Pterostichus hypogeus Barr, 1971
Pterostichus iberlovi O.Berlov, 1996
Pterostichus idahoae Csiki, 1930
Pterostichus ienishteai Nitzu, 1988
Pterostichus ignavus (Tschitscherine, 1897)
Pterostichus ikizderensis Hovorka & Skoupy, 2007
Pterostichus ikukoae Morita, 2007
Pterostichus illigeri (Panzer, 1803)
Pterostichus illustris Leconte, 1851
Pterostichus imasakai Kasahara & Ohtani, 1988
Pterostichus imitatus Morvan, 1978
Pterostichus immarginatus Straneo, 1977
Pterostichus impressicollis (Fairmaire & Laboulbène, 1854)
Pterostichus impressus (Fairmaire & Laboulbène, 1854)
Pterostichus imurai Morita, 2007
Pterostichus inanis G.Horn, 1891
Pterostichus incommodus Schaum, 1858
Pterostichus inermis Fall, 1901
Pterostichus infaustus Gistel, 1857
Pterostichus infernalis Hatch, 1936
Pterostichus infimus Chaudoir, 1868
Pterostichus ingens (Casey, 1918)
Pterostichus ingusha (Lutshnik, 1928)
Pterostichus ingushus (Lutshnik, 1928)
Pterostichus innshanensis Jedlicka, 1960
Pterostichus inopinatus Lutshnik, 1933
Pterostichus inopinus (Casey, 1918)
Pterostichus insidatrix (Piochard de la Brûlerie, 1872)
Pterostichus insidiatrix (Brulerie, 1872)
Pterostichus insidiosus Fairmaire, 1866
Pterostichus insignicollis (Tschitscherine, 1893)
Pterostichus intectus (Casey, 1918)
Pterostichus interruptus (Dejean, 1828)
Pterostichus irideus Sciaky, 1994
Pterostichus iripennis Chaudoir, 1868
Pterostichus isabellae Leconte, 1851
Pterostichus ishiharai Ishida & Shibata, 1961
Pterostichus ishii Morita; Kurosa & Mori, 2009
Pterostichus ishikawai Nemoto, 1988
Pterostichus ishikawaioides Sasakawa; Kim & Kubota, 2008
Pterostichus ishizuchiensis Kasahara, 1985
Pterostichus ishizukai Kasahara, 1995
Pterostichus isolatus Sasakawa; Kim & Kubota, 2008
Pterostichus isumiensis Kasahara & Saito, 1997
Pterostichus itoi Kasahara, 1986
Pterostichus iwakiensis Sasakawa, 2009
Pterostichus ixion (Tschitscherine, 1902)
Pterostichus jacobinus Casey, 1913
Pterostichus jacobsoni Poppius, 1906
Pterostichus jaechi Kirschenhofer, 1997
Pterostichus jaechianus Sciaky & Facchini, 2003
Pterostichus jakobsonianus Lutshnik, 1928
Pterostichus jaljaleensis J.Schmidt, 2009
Pterostichus janatai Sciaky &, Wrase, 1997
Pterostichus janbritoi Morvan, 1994
Pterostichus jani Sciaky & Facchini, 2003
Pterostichus jankowskyi (Tschitscherine, 1897)
Pterostichus janoi Jedlicka, 1952
Pterostichus japonicus (Motschulsky, 1860)
Pterostichus jarkendis Jedlicka, 1965
Pterostichus jelepus Andrewes, 1932
Pterostichus jenisseicus Csiki, 1930
Pterostichus jiricola Sasakawa; Kim & Kubota, 2008
Pterostichus jogaesanensis Lafer; Paik & Park, 1996
Pterostichus johnsoni Ulke, 1889
Pterostichus josephi Csiki, 1930
Pterostichus juga J.schmidt, 2005
Pterostichus jugicola (Lutshnik, 1916)
Pterostichus jugivagus (Tschitscherine, 1898)
Pterostichus jungens (Tschitscherine, 1893)
Pterostichus jureceki (Jedlicka, 1936)
Pterostichus jurecekianus Maran, 1940
Pterostichus jurinei (Panzer, 1803)
Pterostichus justusii W.Redtenbacher, 1842
Pterostichus kadjaranze Jedlicka, 1947
Pterostichus kadleci Dvorak, 1995
Pterostichus kadoudali Morvan, 1981
Pterostichus kajimurai Habu & Tanaka, 1957
Pterostichus kalabi Sciaky, 1994
Pterostichus kalhys (Motschulsky, 1866)
Pterostichus kalo J.Schmidt, 2012
Pterostichus kambaiti (Andrewes, 1947)
Pterostichus kamtschaticus Motschulsky, 1860
Pterostichus kangchenjunga J.Schmidt, 2012
Pterostichus kaniei Morita, 2005
Pterostichus kaninensis Poppius, 1906
Pterostichus kansanus (Casey, 1918)
Pterostichus karasawai Tanaka, 1958
Pterostichus karateghinicus Mikhailov, 1972
Pterostichus kataevi Kryzhanovskij, 1989
Pterostichus katashinensis Habu, 1958
Pterostichus kavanaughi Shilenkov, 2000
Pterostichus kaynashensis Schweiger, 1967
Pterostichus keltiekus Morvan, 1981
Pterostichus kerzhneri Lafer, 1983
Pterostichus kiangsu Jedlicka, 1965
Pterostichus kiiensis Morita & Ohkura, 1988
Pterostichus kimurai Morita, 1994
Pterostichus kiritshenkoi (Lutshnik, 1928)
Pterostichus kirschenblatti Kryzhanovskij, 1988
Pterostichus kitakamisanus Sasakawa, 2005
Pterostichus klapperichi Jedlicka, 1956
Pterostichus kleinfeldi Straneo, 1982
Pterostichus kleinfeldianus Sciaky & Wrase, 1997
Pterostichus koenigi (Reitter, 1887)
Pterostichus koenigianus (Tschitscherine, 1890)
Pterostichus koheii Nakane, 1963
Pterostichus kokeilii L.Miller, 1850
Pterostichus kolosovi Lutshnik, 1928
Pterostichus kolymensis Erjiomin, 1998
Pterostichus komalus (Jedlicka, 1936)
Pterostichus komiyai Morita, 2007
Pterostichus kongosanus Nakane, 1963
Pterostichus kopetzi J.Schmidt, 2005
Pterostichus korgei Jedlicka, 1964
Pterostichus korolkowi (Tschitscherine, 1901)
Pterostichus kosakai Morita, 1998
Pterostichus kotzebuei Ball, 1962
Pterostichus krali Sciaky, 1997
Pterostichus krasnopolensis Kirschenhofer, 1987
Pterostichus krulikovskyi (Lutshnik, 1933)
Pterostichus kubanicus Kirschenhofer, 1987
Pterostichus kucerai Sciaky, 1997
Pterostichus kultianus Jedlicka, 1946
Pterostichus kuraicus Shilenkov, 2000
Pterostichus kurentzovi Lafer, 1979
Pterostichus kurikomasanus Sasakawa, 2005
Pterostichus kurosai Tanaka, 1958
Pterostichus kurosawai Tanaka, 1958
Pterostichus kuwolsanensis Li & Zhang, 2014
Pterostichus kvirensis Belousov, 1991
Pterostichus kyushuensis Habu, 1955
Pterostichus laborans Casey, 1913
Pterostichus labzuki Lafer, 1979
Pterostichus lacertus Casey, 1913
Pterostichus lachrymosus (Newman, 1838)
Pterostichus lacunosus (Chaudoir, 1844)
Pterostichus laevigatus Scudder, 1890
Pterostichus laevipunctatus Tschitscherine, 1889
Pterostichus laferi O. Berlov & E. Berlov, 1996
Pterostichus lama (Ménétriés, 1843)
Pterostichus lamuticus Poppius, 1906
Pterostichus lanei Vandyke, 1925
Pterostichus lanista (Tschitscherine, 1898)
Pterostichus larisae Sundukov, 2013
Pterostichus lasserrei (Dejean, 1828)
Pterostichus lassulus (Casey, 1920)
Pterostichus latemarginatus (Straneo, 1936)
Pterostichus laticollis (Motschulsky, 1844)
Pterostichus latifianus Apfelbeck, 1906
Pterostichus latistylis Tanaka, 1958
Pterostichus latitemporis Sciaky & Wrase, 1997
Pterostichus latiusculus Chaudoir, 1868
Pterostichus lattini Labonte, 2006
Pterostichus ledenikensis (Knirsch, 1925)
Pterostichus leica J.Schmidt, 2012
Pterostichus leonisi Apfelbeck, 1904
Pterostichus leptis Bates, 1873
Pterostichus lesticoides (Straneo, 1939)
Pterostichus letensis Habu, 1973
Pterostichus levadensis Lafer, 1979
Pterostichus leviculus Andrewes, 1933
Pterostichus licenti Jedlicka, 1939
Pterostichus liciniformis Csiki, 1930
Pterostichus lidara Andrewes, 1937
Pterostichus lidarus (Andrewes, 1937)
Pterostichus lineatopunctatus L.Miller, 1850
Pterostichus lingshanus Sciaky &, Wrase, 1997
Pterostichus liodactylus (Tschitscherine, 1898)
Pterostichus lobatus Hacker, 1968
Pterostichus lodosi Heinz, 1977
Pterostichus lolo Bergdahl, 2011
Pterostichus lombardus K.Daniel, 1901
Pterostichus longinquus Bates, 1873
Pterostichus longipennis Straneo, 1942
Pterostichus longipes Poppius, 1906
Pterostichus longissimus Bates, 1882
Pterostichus longiusculus (J.Sahlberg, 1880)
Pterostichus lubricus LeConte, 1853
Pterostichus lucidus Motschulsky, 1844
Pterostichus luctuosus (Dejean, 1828)
Pterostichus lugi Morvan, 1982
Pterostichus lukjanovitshi Lutshnik, 1927
Pterostichus lumensis Apfelbeck, 1905
Pterostichus luscus (Casey, 1918)
Pterostichus lustrans Leconte, 1851
Pterostichus lutschniki Jedlicka, 1962
Pterostichus lutshnikianus Bogachev & Kurnakov, 1958
Pterostichus lyroderus (Chaudoir, 1846)
Pterostichus macedonicus Apfelbeck, 1918
Pterostichus macer (Marsham, 1802)
Pterostichus machulkai Jedlicka, 1931
Pterostichus macrocephalus Habu, 1955
Pterostichus macrogenys Bates, 1883
Pterostichus macrothorax Poppius, 1906
Pterostichus maderi Jedlicka, 1938
Pterostichus madidus (Fabricius, 1775)
Pterostichus maeklini (Poppius, 1906)
Pterostichus magoides (Straneo, 1937)
Pterostichus magus (Mannerheim, 1825)
Pterostichus maichensis Lafer, 1979
Pterostichus makalu J.Schmidt, 2012
Pterostichus makolskii J.Muller, 1933
Pterostichus malaisei Jedlicka, 1965
Pterostichus malissorum Apfelbeck, 1905
Pterostichus malkini (Hatch, 1953)
Pterostichus mancus (Leconte, 1852)
Pterostichus mandibularoides Ball, 1966
Pterostichus mandzhuricus Lutshnik, 1916
Pterostichus mannerheimii (Dejean, 1831)
Pterostichus marani Jedlicka, 1967
Pterostichus marginatus Matsumura, 1911
Pterostichus marginellus Sasakawa; Kim & Kim, 2006
Pterostichus mariae (Lutshnik, 1921)
Pterostichus marinensis Hacker, 1968
Pterostichus masahiroi Kasahara, 1988
Pterostichus masatakai Morita, 2007
Pterostichus masidai Ishida, 1959
Pterostichus masumotoi Tanaka; Morita & Suga, 1987
Pterostichus matchai Jedlicka, 1930
Pterostichus matsumurai Habu, 1973
Pterostichus mattolensis Hacker, 1968
Pterostichus matveichuki O.Berlov, 1996
Pterostichus maurusiacus Mannerheim, 1825
Pterostichus maximus Tschitscherine, 1889
Pterostichus mediocris Dvorak, 1995
Pterostichus medvedevi Kryzhanovskij & Mikhailov, 1972
Pterostichus megacephalus Sciaky & Allegro, 2013
Pterostichus megaloderus Sciaky, 1994
Pterostichus meisteri Reitter, 1885
Pterostichus melanarius (Illiger, 1798)
Pterostichus melanodes (Chaudoir, 1878)
Pterostichus melas (Creutzer, 1799)
Pterostichus mellyi (Gebler, 1843)
Pterostichus menetriesii Leconte, 1873
Pterostichus mercedianus (Casey, 1918)
Pterostichus merkli (J.Frivaldszky, 1879)
Pterostichus merklii (J.Frivaldszky, 1879)
Pterostichus meskheticus Belousov, 1991
Pterostichus metallifer Gistel, 1857
Pterostichus mewakholensis J.Schmidt, 2012
Pterostichus meyeri Jedlicka, 1934
Pterostichus micans Heer, 1838
Pterostichus michailovi Wrase, 1992
Pterostichus microbus Sciaky & Facchini, 2003
Pterostichus microcephalus (Motschulsky, 1860)
Pterostichus microphthalmus Delarouzée, 1857
Pterostichus microps Heyden, 1887
Pterostichus middendorffi J.R. Sahlberg, 1875
Pterostichus miehei J.Schmidt, 2005
Pterostichus migliaccioi Straneo, 1982
Pterostichus militaris (Tschitscherine, 1893)
Pterostichus mimus Casey, 1913
Pterostichus minax (Oustalet, 1874)
Pterostichus ming Sciaky &, Wrase, 1997
Pterostichus minor (Gyllenhal, 1827)
Pterostichus minutulus Heer, 1862
Pterostichus mirabilis Jedlicka, 1939
Pterostichus mirificus Bates, 1883
Pterostichus miroshnikovi Zamotajlov, 1991
Pterostichus miroslavi Sciaky &, Wrase, 1997
Pterostichus mirus (Tschitscherine, 1894)
Pterostichus miscellus Casey, 1913
Pterostichus mitoyamanus Tanaka, 1971
Pterostichus miyamai Kasahara & Y.Ito, 1987
Pterostichus mizunoyai Morita, 2007
Pterostichus modicellus Tschitscherine, 1897
Pterostichus moestus (Say, 1823)
Pterostichus molopsoides Jedlicka, 1934
Pterostichus monostigma (Tschitscherine, 1898)
Pterostichus montanus (Motschulsky, 1844)
Pterostichus monticoloides Shilenkov, 1995
Pterostichus montigena (Tschitscherine, 1898)
Pterostichus montisdeorum Schweiger, 1967
Pterostichus morawitzianus (Lutshnik, 1922)
Pterostichus morio (Duftschmid, 1812)
Pterostichus morionides (Chaudoir, 1868)
Pterostichus mosaicus Sasakawa, 2005
Pterostichus mucronatus Straneo, 1955
Pterostichus muehlfeldii (Duftschmid, 1812)
Pterostichus muelleri (Straneo, 1936)
Pterostichus muellermotzfeldi Wrase & J.Schmidt, 2006
Pterostichus mujahedeeni Savich, 1999
Pterostichus mukdenensis Breit, 1933
Pterostichus mulensis Sciaky, 1997
Pterostichus multinodosus Sasakawa; Kim & Kim, 2006
Pterostichus multipunctatus (Dejean, 1828)
Pterostichus multiseta (Straneo, 1984)
Pterostichus mundatus Jedlicka, 1940
Pterostichus mundus Jedlicka, 1938
Pterostichus musashiensis Kasahara, 1993
Pterostichus mutoides Bousquet, 1992
Pterostichus mutus (Say, 1823)
Pterostichus nadari (Vuillefroy, 1893)
Pterostichus nakanei Straneo, 1955
Pterostichus namrum Jedlicka, 1963
Pterostichus naokii Morita, 2003
Pterostichus napaea Kasahara, 1988
Pterostichus nasuensis Morita, 2007
Pterostichus nasui Kasahara, 1993
Pterostichus necessarius (Tschitscherine, 1894)
Pterostichus neglectus A.Morawitz, 1862
Pterostichus negligens Sturm, 1824
Pterostichus neilgaimani Chaladze, Kalatozishvili, Janiashvili, & Bakuradze 2017[6]
Pterostichus nemoralis Graells, 1851
Pterostichus neobrunneus Lindroth, 1966
Pterostichus nepalensis Straneo, 1977
Pterostichus nicaeensis A. Villa & C.B. Villa, 1835
Pterostichus nigellatus Kirschenhofer, 1991
Pterostichus nigellus A.Morawitz, 1862
Pterostichus niger (Schaller, 1783)
Pterostichus nigripalpis Poppius, 1906
Pterostichus nigrita (Paykull, 1790)
Pterostichus nigrocaeruleus Vandyke, 1925
Pterostichus nigrocoerulus
Pterostichus nikodymi Dvorak, 1995
Pterostichus nimbatidius Chaudoir, 1878
Pterostichus nishidai Nakane, 1989
Pterostichus nishiyamai Kasahara, 1986
Pterostichus nivalis R.F. Sahlberg, 1844
Pterostichus nivicola (Ménétriés, 1832)
Pterostichus noburai Morita, 2008
Pterostichus nodicornis (Fairmaire & Laboulbène, 1854)
Pterostichus noguchii Bates, 1873
Pterostichus nordqvisti J.R. Sahlberg, 1887
Pterostichus novotnyorum Vysoky, 1981
Pterostichus novus Straneo, 1944
Pterostichus nowitzkii (Tschitscherine, 1899)
Pterostichus obesulus Leconte, 1873
Pterostichus oblongiusculus Motschulsky, 1850
Pterostichus oblongoparallelus Maran, 1932
Pterostichus oblongopunctatus (Fabricius, 1787)
Pterostichus obsidianus Casey, 1913
Pterostichus occultus Casey, 1913
Pterostichus ochoticus R.F. Sahlberg, 1844
Pterostichus ochraceus Gautier des Cottes, 1868
Pterostichus ochsi Fagniez, 1921
Pterostichus odontocnemis G.Müller, 1932
Pterostichus odvarkai Dvorak, 1995
Pterostichus oenotrius Ravizza, 1975
Pterostichus ogaensis Morita, 1995
Pterostichus ohbayashii Ishida, 1968
Pterostichus ohdaisanus Nakane, 1963
Pterostichus ohkawai Morita, 2010
Pterostichus ohkurai Morita, 1996
Pterostichus ohsawacavus Sasakawa, 2005
Pterostichus okiensis Nakane, 1989
Pterostichus okutamae Tanaka, 1963
Pterostichus olafi J.Schmidt, 2005
Pterostichus olegi Belousov, 1991
Pterostichus omogoensis Nakane, 1972
Pterostichus ompoensis Jedlicka, 1932
Pterostichus opacipennis Jedlicka, 1934
Pterostichus ophryoderus (Chaudoir, 1878)
Pterostichus ordinarius Casey, 1913
Pterostichus ordinatus (Fischer, 1823)
Pterostichus orduensis (Straneo, 1988)
Pterostichus oregonus Leconte, 1861
Pterostichus oreophilus (Tschitscherine, 1898)
Pterostichus orestes (Jedlicka, 1936)
Pterostichus orientalis (Motschulsky, 1844)
Pterostichus orion (Tschitscherine, 1901)
Pterostichus orionis Jedlicka, 1962
Pterostichus orman Jedlicka, 1963
Pterostichus orobius Fairmaire, 1861
Pterostichus oscitans Tschitscherine, 1900
Pterostichus osseticus Kirschenhofer, 1987
Pterostichus ottomanus Apfelbeck, 1908
Pterostichus ovalipennis Casey, 1913
Pterostichus ovaliphallus Sasakawa, 2005
Pterostichus ovoideus (Sturm, 1824)
Pterostichus pachinus Bates, 1883
Pterostichus palekhai E. Berlov & O. Berlov, 1996
Pterostichus palgongsanus Nemoto, 1988
Pterostichus palmi Schaeffer, 1910
Pterostichus panticulatus Casey, 1913
Pterostichus pantomus Jedlicka, 1958
Pterostichus panzeri (Panzer, 1803)
Pterostichus paralleloides Kirschenhofer, 1987
Pterostichus parasimilis G.E. Ball, 1962
Pterostichus parkwon Davies, 2004
Pterostichus parnassius Schaum, 1859
Pterostichus parviceps Poppius, 1906
Pterostichus parvicollis Sciaky &, Wrase, 1997
Pterostichus pasanauricus Kirschenhofer, 1987
Pterostichus patruelis (Dejean, 1831)
Pterostichus paulinoi (Vuillefroy, 1868)
Pterostichus pavani (Straneo, 1988)
Pterostichus pedemontanus Ganglbauer, 1891
Pterostichus peilingi Jedlicka, 1937
Pterostichus pemphredo Will, 2007
Pterostichus peninsularis Park & Kwon, 1996
Pterostichus pensylvanicus Leconte, 1873
Pterostichus pentheri Apfelbeck, 1918
Pterostichus percontator Reitter, 1887
Pterostichus pergracilis (Casey, 1920)
Pterostichus perhoplites J.Schmidt & Tian, 2011
Pterostichus perisi Novoa, 1979
Pterostichus perlamatus Straneo, 1989
Pterostichus perlutus Jedlicka, 1938
Pterostichus permundus (Say, 1830)
Pterostichus pertinax (Tschitscherine, 1895)
Pterostichus petulans Jedlicka, 1938
Pterostichus pfizenmayeri Poppius, 1906
Pterostichus phaeopus (Sainte-Claire Deville, 1902)
Pterostichus phaeus (Lutshnik, 1922)
Pterostichus phungaraziensis Wrase & J.Schmidt, 2006
Pterostichus piceolus (Chaudoir, 1850)
Pterostichus picipes Fischer von Waldheim, 1829
Pterostichus pilifer (Bates, 1878)
Pterostichus piliferus (Bates, 1878)
Pterostichus pilosus (Host, 1789)
Pterostichus pinguedineus Eschscholtz, 1823
Pterostichus piniphilus Gistel, 1857
Pterostichus pithyusicus Lutshnik, 1933
Pterostichus placerensis (Casey, 1918)
Pterostichus planaticollis Kirschenhofer, 1987
Pterostichus planctus Leconte, 1852
Pterostichus planicola (Tschitscherine, 1899)
Pterostichus planiusculus (Chaudoir, 1859)
Pterostichus planus J.R. Sahlberg, 1887
Pterostichus platyderus Chaudoir, 1850
Pterostichus platyops Sciaky, 1997
Pterostichus plesiomorphus Nemoto, 1989
Pterostichus plutonicus Casey, 1913
Pterostichus poecilma Andrewes, 1937
Pterostichus pohnerti Jedlicka, 1934
Pterostichus polygenus Bates, 1883
Pterostichus ponticus Kirschenhofer, 1987
Pterostichus poppiusi (Semenov, 1906)
Pterostichus poppiusianus Jakobson, 1907
Pterostichus porcellus Kurnakov, 1962
Pterostichus potanini Tschitscherine, 1889
Pterostichus praedo (Tschitscherine, 1901)
Pterostichus praetermissus (Chaudoir, 1868)
Pterostichus pratti Bates, 1890
Pterostichus prattii Bates, 1890
Pterostichus primus Darlington, 1932
Pterostichus probus Park & Kwon, 1996
Pterostichus procax A.Morawitz, 1862
Pterostichus procephalus Bates, 1873
Pterostichus procerulus Heyden, 1880
Pterostichus properans Chaudoir, 1868
Pterostichus protensiformis (Casey, 1924)
Pterostichus protensipennis (Casey, 1918)
Pterostichus protractus Leconte, 1860
Pterostichus provincialis (Oustalet, 1874)
Pterostichus przewalskyi Tschitscherine, 1888
Pterostichus pseodostuxbergi Poppius, 1906
Pterostichus pseudoastutus Hovorka & Skoupy, 2007
Pterostichus pseudobarbarae Sciaky & Facchini, 2003
Pterostichus pseudodiversus Shi & Sciaky, 2013
Pterostichus pseudoharmandi Morvan, 1981
Pterostichus pseudojugivagus J.Schmidt & Tian, 2011
Pterostichus pseudopachinus Nakane, 1963
Pterostichus pseudopedius Reitter, 1887
Pterostichus pseudopercus Reitter, 1889
Pterostichus pseudopilifer (Straneo, 1957)
Pterostichus pseudoplatyderus Sciaky, 1994
Pterostichus pseudoplatysma Straneo, 1982
Pterostichus pseudorotundus Sciaky & Facchini, 2003
Pterostichus pseudosinensis Sciaky & Facchini, 2003
Pterostichus pseudostuxbergi (Poppius, 1906)
Pterostichus pulchellus Faldermann, 1836
Pterostichus pulcher Sciaky & Allegro, 2013
Pterostichus pumilio Dejean, 1828
Pterostichus pumilus Casey, 1913
Pterostichus pumpellyi Scudder, 1900
Pterostichus punctatissimus (Randall, 1838)
Pterostichus punctatus L. Redtenbacher, 1843
Pterostichus punctibasalis N.Ito, 2012
Pterostichus punctiger J.R. Sahlberg, 1880
Pterostichus punctisternus Straneo, 1989
Pterostichus punctiventris (Chaudoir, 1878)
Pterostichus pusillus Dejean, 1828
Pterostichus putus Casey, 1913
Pterostichus quadraticollis Chaudoir, 1846
Pterostichus quadrifoveolatus Letzner, 1852
Pterostichus quadriimpressus Straneo, 1984
Pterostichus quelpartensis Habu, 1985
Pterostichus raptor (Tschitscherine, 1901)
Pterostichus raraensis Morvan, 1980
Pterostichus rasilis Park & Kwon, 1996
Pterostichus regeli (Tschitscherine, 1894)
Pterostichus reiseri Ganglbauer, 1889
Pterostichus reissi G.Müller, 1931
Pterostichus relictus (Newman, 1838)
Pterostichus rengensis Morita, 2010
Pterostichus restrictus (Casey, 1918)
Pterostichus reuteri Wrase & Schmidt, 2006
Pterostichus rhaeticus Heer, 1837
Pterostichus rhanis (Tschitscherine, 1902)
Pterostichus rhilensis Rottenberg, 1874
Pterostichus rifensis Antoine, 1933
Pterostichus riffensis Antoine, 1933
Pterostichus riparius (Dejean, 1828)
Pterostichus ripensis (Motschulsky, 1866)
Pterostichus robustistylis Sasakawa, 2005
Pterostichus rodionovi Shilenkov 2000 RU (Chita Area)
Pterostichus rolex Morvan, 1995
Pterostichus rostratus (Newman, 1838)
Pterostichus rothi (Hatch, 1951)
Pterostichus rotundangulus A.Morawitz, 1862
Pterostichus rotundus Sciaky, 1994
Pterostichus rousi Kirschenhofer, 1982
Pterostichus rousianus Kirschenhofer, 1987
Pterostichus rousselli Colas, 1963
Pterostichus rubripalpis Csiki, 1930
Pterostichus rudestriatus (Reitter, 1883)
Pterostichus ruffoi Sciaky, 1984
Pterostichus rufipes (Dejean, 1828)
Pterostichus rufitarsis (Dejean, 1828)
Pterostichus rufonitens Fairmaire, 1866
Pterostichus rufopiceus Heyden, 1890
Pterostichus rugiceps (Straneo, 1984)
Pterostichus rugosiceps J.Schmidt, 2012
Pterostichus rugosipennis Jedlicka, 1932
Pterostichus rugosus (Gebler, 1823)
Pterostichus rutenus (Jeannel, 1942)
Pterostichus rutilans (Dejean, 1828)
Pterostichus ryomoensis Morita & Suda, 2007
Pterostichus sacheri I.Frivaldszky von Frivald, 1865
Pterostichus sachtlebeni Jedlicka, 1962
Pterostichus safonovae Budarin, 1995
Pterostichus sagarmatha J.Schmidt, 2012
Pterostichus sahlbergi (Tschitscherine, 1894)
Pterostichus sakagamii Morita, 2001
Pterostichus sakuraii Morita, 2008
Pterostichus santostamangi J.Schmidt, 2006
Pterostichus sasajii Morita, 2007
Pterostichus satoi Ishida, 1961
Pterostichus satsumanus Habu, 1958
Pterostichus satunini Tschitscherine, 1903
Pterostichus satyrus Kurnakov, 1962
Pterostichus saueri Sciaky, 1994
Pterostichus saxicola (Tschitscherine, 1899)
Pterostichus sayanus Csiki, 1930
Pterostichus scalptus Sciaky & Wrase, 1997
Pterostichus schaschli (Marseul, 1880)
Pterostichus schmidtii (Chaudoir, 1838)
Pterostichus schneideri Sciaky & Facchini, 2003
Pterostichus schodaicus Kirschenhofer, 1987
Pterostichus schoenherri Faldermann, 1836
Pterostichus schoenmanni Kirschenhofer, 1991
Pterostichus schrettenbrunneri J.Schmidt, 2012
Pterostichus schuelkei Sciaky & Wrase, 1997
Pterostichus scitus Maklin, 1878
Pterostichus sculpturalis N.Ito, 2012
Pterostichus sculptus LeConte, 1853
Pterostichus scurra (Tschitscherine, 1901)
Pterostichus scutellaris Leconte, 1873
Pterostichus scuticollis (Fairmaire, 1889)
Pterostichus sediorcus Sasakawa & Kim, 2006
Pterostichus sejunctus Bates, 1883
Pterostichus sejungendus (Chaudoir, 1868)
Pterostichus selmanni (Duftschmid, 1812)
Pterostichus semenovi Tschitscherine, 1888
Pterostichus semirugosus (Andrewes, 1947)
Pterostichus septemtrionalis Sciaky & Facchini, 2003
Pterostichus septentrionis (Chaudoir, 1868)
Pterostichus serbicus Apfelbeck, 1899
Pterostichus seriatus (Chaudoir, 1850)
Pterostichus serratipes Chaudoir, 1878
Pterostichus serripes (Leconte, 1875)
Pterostichus seticeps (Straneo, 1984)
Pterostichus setipes (Tschitscherine, 1898)
Pterostichus setosicus Davies, 2004
Pterostichus setosus Hatch, 1951
Pterostichus seunglaki Park & Kwon, 1996
Pterostichus seungmoi Park & Kwon, 1996
Pterostichus shaovuensis Jedlicka, 1956
Pterostichus sharanus (Straneo, 1984)
Pterostichus shavrini O.Berlov & Anistschenko, 1999
Pterostichus shennongjianus Facchini & Sciaky, 2003
Pterostichus shibatai Ishida, 1961
Pterostichus shiibanus Habu, 1958
Pterostichus shilenkovi Erjiomin & Kabak, 1991
Pterostichus shimizui Morita, 2010
Pterostichus shinbodakensis Sasakawa & H.Ito, 2017
Pterostichus shingarevi Lafer, 1979
Pterostichus shirakamisan Sasakawa, 2009
Pterostichus shirakamisanus Sasakawa, 2005
Pterostichus shiranus (Straneo, 1984)
Pterostichus shirozui Habu, 1985
Pterostichus shojii Sugimura, 2006
Pterostichus shotaroi Morita, 1987
Pterostichus shulli (Hatch, 1949)
Pterostichus sierranus Casey, 1913
Pterostichus similatus Gautier des Cottes, 1870
Pterostichus similis Mannerheim, 1852
Pterostichus simillimus Fairmaire, 1886
Pterostichus sincerus Park & Kwon, 1996
Pterostichus sinensis Jedlicka, 1962
Pterostichus singularis Tschitscherine, 1889
Pterostichus sinicus (Fairmaire, 1897)
Pterostichus sintanus Andrewes, 1927
Pterostichus skoupyi Hovorka & Skoupy, 2007
Pterostichus smetanai Bousquet, 1985
Pterostichus sodalicius Heyden, 1885
Pterostichus sojaki Kirschenhofer, 1987
Pterostichus sojot Shilenkov, 2000
Pterostichus solskyi (Chaudoir, 1878)
Pterostichus songoricus (Motschulsky, 1845)
Pterostichus soperi Ball, 1966
Pterostichus sotkaensis Jedlicka, 1958
Pterostichus spathifer Bousquet, 1992
Pterostichus sphodriformis Bates, 1873
Pterostichus sphodrinus Leconte, 1863
Pterostichus spiculifer Bates, 1883
Pterostichus spinolae (Dejean, 1828)
Pterostichus sponsor Casey, 1913
Pterostichus spornyi Sundukov, 2013
Pterostichus spraguei Leconte, 1873
Pterostichus ssemenovi Tschitscherine, 1888
Pterostichus stagnatilis Gistel, 1857
Pterostichus stantonensis Ball, 1966
Pterostichus stapedius Hacker, 1968
Pterostichus starcki Heyden, 1885
Pterostichus starckianus Reitter, 1887
Pterostichus staveni Hovorka & Skoupy, 2007
Pterostichus sterbai Jedlicka, 1934
Pterostichus stictopleurus (Fairmaire, 1889)
Pterostichus stoeckleini (Straneo, 1941)
Pterostichus stomoides Chaudoir, 1868
Pterostichus straneellus Jedlicka, 1938
Pterostichus strasseri Reitter, 1898
Pterostichus strenuus (Panzer, 1796)
Pterostichus stricticollis (Solsky, 1874)
Pterostichus strigipennis N.Ito, 2012
Pterostichus strigosus Sciaky & Wrase, 1997
Pterostichus stygicus (Say, 1823)
Pterostichus styx Andrewes, 1937
Pterostichus subabacipennis N.Ito, 2012
Pterostichus subacutus (Casey, 1918)
Pterostichus subaeneus (Chaudoir, 1850)
Pterostichus subgibbus Motschulsky, 1860
Pterostichus subiasi Ortuno & Zaballos, 1992
Pterostichus subitus Csiki, 1930
Pterostichus sublaevis (J.Sahlberg, 1880)
Pterostichus subovatus (Motschulsky, 1860)
Pterostichus subrugosus Straneo, 1955
Pterostichus subsinuatus Dejean, 1828
Pterostichus subtilissimus Sciaky, 1996
Pterostichus sudai Morita, 1991
Pterostichus sudrei Sainte-Claire Deville, 1922
Pterostichus suffusus Casey, 1913
Pterostichus sugimurai Morita, 2007
Pterostichus sulcitarsis A.Morawitz, 1863
Pterostichus sumondakensis Sasakawa, 2005
Pterostichus sungariensis Lafer, 1979
Pterostichus superciliosus (Say, 1823)
Pterostichus surgens Leconte, 1878
Pterostichus suruganus Straneo, 1979
Pterostichus sutschanensis Jedlicka, 1962
Pterostichus svanicus (Lutshnik, 1922)
Pterostichus swaneticus (Reitter, 1883)
Pterostichus syleus Kirschenhofer, 1997
Pterostichus symmetricus Straneo, 1955
Pterostichus szekessyianus Sciaky, 1996
Pterostichus szetschuanensis Tschitscherine, 1889
Pterostichus tachongi Jedlicka, 1936
Pterostichus taebaegsanus Nemoto, 1988
Pterostichus taeniolus Bates, 1882
Pterostichus tahirai Kasahara, 1992
Pterostichus tahoensis Casey, 1913
Pterostichus takadateyamanus Sasakawa, 2009
Pterostichus takahashii Ishida, 1958
Pterostichus takaosanus Habu, 1958
Pterostichus taksonyis Csiki, 1930
Pterostichus talibani Savich, 1999
Pterostichus tamarae Wrase & Kirschenhofer, 1991
Pterostichus tamsii (Dejean, 1831)
Pterostichus tanakai Ishida, 1964
Pterostichus tanakaorum Morita & Ohkura, 1988
Pterostichus tanjae Hovorka & Skoupy, 2007
Pterostichus tantillus Fairmaire, 1888
Pterostichus taoi Kasahara, 1993
Pterostichus taradakensis Kasahara & Ohtani, 1988
Pterostichus tareumiut G. E. Ball, 1962
Pterostichus tarsalis Leconte, 1873
Pterostichus taskylensis Shilenkov, 2000
Pterostichus tateishiyamanus Sasakawa & H.Ito, 2017
Pterostichus tatianae E.Berlov, 1996
Pterostichus tatricus Kult, 1947
Pterostichus tenenbaumianus Jedlicka, 1930
Pterostichus tenuimarginatus Chaudoir, 1868
Pterostichus tenuis (Casey, 1924)
Pterostichus teretis Park & Kwon, 1996
Pterostichus testaceus (Vandyke, 1926)
Pterostichus thanglaensis J.Schmidt, 2012
Pterostichus theeli Maklin, 1878
Pterostichus thorectes Bates, 1873
Pterostichus thorectoides Jedlicka, 1958
Pterostichus thunbergi A.Morawitz, 1862
Pterostichus tichomirovi Erjiomin, 1990
Pterostichus tienmushanus Sciaky, 1997
Pterostichus tiliaceoradix Ball, 1962
Pterostichus todai Morita & Kanie, 1997
Pterostichus tokejii Yoshida & Tanaka, 1960
Pterostichus tokmakovae Sundukov, 2013
Pterostichus tokui Morita, 2007
Pterostichus toledanoi Facchini & Sciaky, 2003
Pterostichus tomensis (Gebler, 1847)
Pterostichus tonkinensis Straneo, 1980
Pterostichus tosanus Kasahara & Y.ito, 1997
Pterostichus tottoriensis Morita, 2003
Pterostichus toyodai Morita & Kurosa, 1998
Pterostichus transversalis (Duftschmid, 1812)
Pterostichus triangularis Sciaky & Facchini, 2003
Pterostichus trinarius (Casey, 1918)
Pterostichus trinitensis Hacker, 1968
Pterostichus triseriatus (Gebler, 1847)
Pterostichus tristis (Dejean, 1828)
Pterostichus troglophilus Mateu, 1997
Pterostichus tropicalis Bates, 1882
Pterostichus trubilini Zamotajlov, 1997
Pterostichus truncatus (Dejean, 1828)
Pterostichus tsagerkericus (Lutshnik, 1928)
Pterostichus tschitscherinianus (Jakobson, 1907)
Pterostichus tsukubasanus Kasahara, 1988
Pterostichus tsurugaensis Ishida, 1959
Pterostichus tsurugiyamanus Habu, 1959
Pterostichus tuberculiger (Tschitscherine, 1897)
Pterostichus tuberculofemoratus Hatch, 1936
Pterostichus tuberifer Sasakawa, 2006
Pterostichus tulobalu Schmidt, 2009
Pterostichus tumescens Leconte, 1863
Pterostichus tundrae (Tschitscherine, 1894)
Pterostichus tunkinensis Shilenkov, 2000
Pterostichus turanensis Jedlicka, 1959
Pterostichus turcomannicus Motschulsky, 1850
Pterostichus uchiyamai Morita, 1987
Pterostichus udokanensis Shilenkov, 2000
Pterostichus uedaorum Morita & Hirasawa, 1996
Pterostichus uenoi Straneo, 1955
Pterostichus ulubeyensis Kirschenhofer, 1981
Pterostichus umbralis Gistel, 1857
Pterostichus umbricola Gistel, 1857
Pterostichus unctulatus Duftschmid, 1812
Pterostichus uralensis Motschulsky, 1850
Pterostichus urengaicus Jurecek, 1924
Pterostichus ussuriensis (Tschitscherine, 1897)
Pterostichus vagepunctatus Heer, 1837
Pterostichus validiceps Reitter, 1887
Pterostichus validior Tschitscherine, 1889
Pterostichus validus (Dejean, 1828)
Pterostichus vandykei Schaeffer, 1910
Pterostichus variabilis (Ménétriés, 1832)
Pterostichus variicornis Sasakawa; Kim & Kim, 2009
Pterostichus variolatus (Dejean, 1828)
Pterostichus variseta (Straneo, 1984)
Pterostichus vecors (Tschitscherine, 1897)
Pterostichus ventralis (Say, 1823)
Pterostichus ventricosus Eschscholtz, 1823
Pterostichus vermiculosus (Ménétriés, 1851)
Pterostichus vernalis Panzer, 1796
Pterostichus vexatus Bousquet, 1985
Pterostichus vicinus Mannerheim, 1843
Pterostichus violaceotinctus Straneo, 1953
Pterostichus virescens (Gebler, 1833)
Pterostichus viridellus Straneo, 1984
Pterostichus vladivostokensis Lafer, 1979
Pterostichus vseteckai (Maran, 1944)
Pterostichus vulgaris (Linnaeus, 1758)
Pterostichus walcotti Scudder, 1900
Pterostichus walteri Reitter, 1883
Pterostichus watanabei Nakane, 1960
Pterostichus weigeli J.Schmidt, 2005
Pterostichus weiperti J.Schmidt, 2005
Pterostichus wenxianensis Allegro & Sciaky, 2010
Pterostichus wittmeri Morvan, 1978
Pterostichus woodi Ball and Currie, 1997
Pterostichus woongbii Park & Kwon, 1996
Pterostichus woronowi (Lutshnik, 1916)
Pterostichus wrasei Sciaky & Facchini, 2003
Pterostichus xatartii (Dejean, 1828)
Pterostichus xilingensis Allegro & Sciaky, 2010
Pterostichus yahikosanus Sasakawa, 2009
Pterostichus yakushimanus Nakane & Ishida, 1961
Pterostichus yamajii Kasahara, 1993
Pterostichus yamauchii Morita, 1992
Pterostichus yamizosanus Sasakawa, 2005
Pterostichus yanoi Jedlicka, 1953
Pterostichus yatsuensis Straneo, 1955
Pterostichus ybousqueti O.Berlov, 1999
Pterostichus yokohamae Nakane & Straneo in Nakane, 1979
Pterostichus yorikoae Sugimura, 2006
Pterostichus yoritomus Bates, 1873
Pterostichus yosemitensis Hacker, 1968
Pterostichus yoshidai Kasahara, 1985
Pterostichus yoshikawai Ishida, 1959
Pterostichus yoshizawai Morita, 2010
Pterostichus yulongshanensis Sciaky, 1997
Pterostichus yunnanensis Jedlicka, 1934
Pterostichus yvanii (Dejean, 1828)
Pterostichus zamotajlovi Belousov, 1991
Pterostichus zhejiangensis Kirschenhofer, 1997
Pterostichus ziegleri (Duftschmid, 1812)
Pterostichus zoiai Sciaky, 1996
Pterostichus zolotarewi Reitter, 1911
Pterostichus zunianus Casey, 1913
Pterostichus zyzzovi O. Berlov & E. Berlov, 1996
References
Bervoets, L; Int Panis, L; Verheyen, R.F. (1994). "Habitat preference and activity period of some carabid species (Coleoptera, Carabidae) in a heathland ecosystem based on a four day sampling period". Bull. Ann. Soc. R. Ent. Belg. 130: 115–118.
"Pterostichus Bonelli, 1810". Carabidae of the World. 2011. Retrieved 4 Jun 2013.
"Pterostichus Genus Information". BugGuide.net. Retrieved 2018-03-06.
"Pterostichus Report". Integrated Taxonomic Information System. Retrieved 2018-03-06.
"Browse Pterostichus". Catalogue of Life. Retrieved 2018-03-06.
Chaladze G, Kalatozishvili L, Janiashvili Z, Bakuradze G (2017). "Pterostichus neilgaimani sp. nov., a new species of ground beetles (Coleoptera: Carabidae) from relict sacred grove in Eastern Georgia". Zootaxa. 4329 (1): 091–095. doi:10.11646/zootaxa.4329.1.6. ISSN 1175-5334. PMID 29242485. Retrieved 4 Oct 2017.
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